After Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009, with species from Uetz, Etzold & Chenna, 1995-2015 [“The Reptile Database”]
<==o Leptotyphlopidae Stejneger, 1892 (matokäärmeet; Slender Blind Snakes) |--o Epictinae Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | |--o Rhinoleptini Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | | |--o Tricheilostoma Jan, 1860 sensu Hedges, 2011 [Guinea Hedges, Adalsteinsson &: Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009, non Guinea Koçak & Kemal, 2008 (Insecta: Hymenoptera)] | | | |-- G. bicolor (Jan 1860) [Leptotyphlops bicolor] | | | |-- G. broadleyi (Wallach & Hahn 1997) [Leptotyphlops broadleyi] | | | |-- G. greenwelli (Wallach & Boundy 2005) [Leptotyphlops greenwelli] | | | `-- G. sundewalli (Jan 1861) [Leptotyphlops sundewalli] | | `--o Rhinoleptus Orejas-Miranda, Roux-Estève & Guibé, 1970 | | |-- R. koniagui (Villiers, 1956) [Typhlops koniagui Villiers, 1956; Leptotyphlops koniagui Guibé, Roux-Estève & Villiers, 1967; Rhinoleptus koniagui Orejas-Miranda, Roux-Estève & Guibé, 1970] (Villiers' blind snake) | | `-- R. parkeri (Broadley, 1999) | `--o Epictini Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | |--o Epictina Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | | |--o Epictia Gray, 1845 [Leptotyphlops Fitzinger, 1843 in partim] | | | |-- E. albipuncta (Burmeister, 1861) | | | |-- E. alfredschmidti (Lehr, Wallach, Köhler & Aguilar, 2002) | | | |-- E. ater (Taylor, 1940) | | | |-- E. australis (Freiberg & Orejes-Miranda, 1968) | | | |-- E. borapeliotes (Vanzolini, 1996) | | | |-- E. clinorostris Arredondo & Zaher, 2010 | | | |-- E. collaris (Hoogmoed, 1977) | | | |-- E. columbi (Klauber, 1939) | | | |-- E. diaplocia (Orejes-Miranda, 1969) | | | |-- E. goudotii (Duméril & Bibron, 1844) | | | |-- E. magnamaculata (Taylor, 1940) | | | |-- E. melanurus (Schmidt, 1943) | | | |-- E. munoai (Orejes-Miranda, 1961) | | | |-- E. nasalis (Taylor, 1940) | | | |-- E. peruviana (Orejes-Miranda, 1969) | | | |-- E. rubrolineata (Werner, 1901) | | | |-- E. rufidorsa (Taylor, 1940) | | | |-- E. signata (Jan, 1861) | | | |-- E. striatula (Smith & Laufe, 1945) | | | |-- E. subcrotilla (Klauber, 1939) | | | |-- E. teaguei (Orejes-Miranda, 1964) | | | |-- E. tenella (Klauber, 1939) | | | |-- E. tesselata (Tschudi, 1845) | | | |-- E. tricolor (Orejes-Miranda & Zug, 1974) | | | |-- E. undecimstriata (Schlegel, 1839) | | | |-- E. unicolor Werner, 1913 | | | `-- E. vellardi (Laurent, 1984) | | `--o Siagonodon Peters, 1881 | | |-- S. acutirostris Pinto & Curcio, 2011 | | |-- S. borrichianus (Degerbøl, 1923) | | |-- S. cupinensis (Bailey & Carvalho, 1946) | | `-- S. septemstriatus (Schneider, 1801) | `--+--o Renina Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | | |--o Rena Baird & Girard, 1853 | | | |-- R. affinis (Boulenger, 1884) | | | |-- R. boettgeri (Werner, 1899) | | | |-- R. bressoni (Taylor, 1939) | | | |-- R. dissectus (Cope, 1896) | | | |-- R. dulcis Baird & Girard, 1853 | | | |-- R. humilis Baird & Girard, 1853 | | | |-- R. maxima (Loveridge, 1932) | | | |-- R. myopica (Garman, 1884) | | | `-- R. unguirostris (Boulenger, 1902) | | `--o Trilepida Hedges, 2011 [Tricheilostoma Jan, 1860 in partim] | | |-- T. anthracina (Bailey, 1946) | | |-- T. brasiliensis (Laurent, 1949) | | |-- T. brevissima (Shreve, 1964) | | |-- T. dimidiatum (Jan, 1861) | | |-- T. dugandi (Dunn, 1944) | | |-- T. fuliginosa (Passos, Caramaschi & Pinto, 2006) | | |-- T. guayaquilensis (Orejes-Miranda & Peters, 1970) | | |-- T. jani Pinto & Fernandes, 2012 | | |-- T. joshuai (Dunn, 1944) | | |-- T. koppesi (Amaral, 1955) | | |-- T. macrolepis (Peters, 1857) | | |-- T. nicefori (Dunn, 1946) | | `-- T. salgueiroi (Amaral, 1955) | `--o Tetracheilostomina Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | |--o Mitophis Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | | |-- M. asbolepis (Thomas, McDiarmid & Thompson, 1985) | | |-- M. calypso (Thomas, McDiarmid & Thompson, 1985) | | |-- M. leptipileptus (Thomas, McDiarmid & Thompson, 1985) | | `-- M. pyrites (Thomas, 1965) | `--o Tetracheilostoma Jan, 1861 | |-- T. bilineatum (Schlegel, 1839) | |-- T. breuili (Hedges, 2008) | |-- T. carlae (Hedges, 2008) | `-- T. dissimilis (Bocage, 1886) `--o Leptotyphlopinae Stejneger, 1892 |?-o Epacrophini Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | `--o Epacrophis Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | |-- E. boulengeri (Boettger, 1913) | |-- E. drewesi (Wallach, 1996) | `-- E. reticulatus (Boulenger, 1906) `--+--o Myriopholini Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | `--o Myriopholis Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 | |-- M. adleri (Hahn & Wallach, 1998) | |-- M. albiventer (Hallermann & Rödel, 1995) | |-- M. algeriensis (Jacquet, 1895) | |-- M. blanfordi (Boulenger, 1890) | |-- M. boueti (Chabanaud, 1917) | |-- M. braccianii (Scortecci, 1929) | |-- M. burii (Boulenger, 1905) | |-- M. cairi (Duméril & Bibron, 1844) | |-- M. erythraeus (Scortecci, 1929) | |-- M. filiformis (Boulenger, 1899) | |-- M. ionidesi (Broadley & Wallach, 2007) | |-- M. longicauda (Peters, 1854) | |-- M. macrorhyncha (Jan, 1860) | |-- M. macrura (Boulenger, 1899) | |-- M. narirostris (Peters, 1867) | |-- M. natatrix (Andersson, 1937) | |-- M. perreti (Roux-Estève, 1979) | |-- M. rouxestevae (Trape & Mane, 2004) | |-- M. wilsoni (Hahn, 1978) | `-- M. yemenicus (Scortecci, 1933) `--o Leptotyphlopini Stejneger, 1892 sensu Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 |--o Leptotyphlops Fitzinger, 1843 [Stenostoma Wagler, 1824 (preocc.); Typhlina Wagler, 1830; Stenostona Cuvier, 1836; Leptotyphlops Fitzinger, 1843; Eucephalus Fitzinger, 1843; Catodon Duméril & Bibron, 1844; Glauconia Gray, 1845; Sabrina Girard, 1857; Ramphostoma Jan, in Jan & Sordelli, 1860; Tricheilostoma Jan, in Jan & Sordelli, 1860; Tetracheilostoma Jan, 1861; Stenostomophis Rochebrune, 1884; Rhamphostoma Boulenger, 1893] (Slender blind snakes, thread snakes) | |-- L. aethiopicus Broadley & Wallach, 2007 | |-- L. conjunctus (Jan, 1861) | |-- L. distanti (Boulenger, 1892) | |-- L. emini (Boulenger, 1890) | |-- L. howelli Broadley & Wallach, 2007 | |-- L. incognitus (Broadley & Watson, 1976) | |-- L. jacobseni Broadley & Broadley, 1999 | |-- L. kafubi (Boulenger, 1919) | |-- L. keniensis Broadley & Wallach, 2007 | |-- L. latirostris (Sternfeld, 1912) | |-- L. macrops (Broadley & Wallach, 1996) | |-- L. mbanjensis Broadley & Wallach, 2007 | |-- L. merkeri (Werner, 1909) | |-- L. nigricans (Schlegel, 1839) | |-- L. nigroterminus Broadley & Wallach, 2007 | |-- L. nursii (Anderson, 1896) | |-- L. pembae Loveridge, 1941 | |-- L. pitmani Broadley & Wallach, 2007 | |-- L. pungwensis Broadley & Wallach, 1997 | |-- L. scutifrons (Peters, 1854) | |-- L. sylvicolus Broadley & Wallach, 1997 | |-- L. tanae (Broadley & amp;Wallach, 2007) | `-- L. telloi Broadley & Watson, 1976 `--o Namibiana Hedges, Adalsteinsson & Branch, in Adalsteinsson, Branch, Trape, Vitt & Hedges, 2009 |-- N. gracilior (Boulenger, 1910) |-- N. labialis (Sternfeld, 1908) |-- N. latifrons (Sternfeld, 1908) |-- N. occidentalis (Fitzsimons, 1962) `-- N. rostrata (Bocage, 1886)
Reference(s):
- Adalsteinsson, S. A., Branch, W. R., Trape, S., Vitt, L. J. & Hedges, S. B., 2009: Molecular phylogeny, classification, and biogeography of snakes of the Family Leptotyphlopidae (Reptilia, Squamata).
–Zootaxa: Vol. 2244, pp. 1-50 - Hedges, S. B., 2011: The type species of the threadsnake genus Tricheilostoma Jan revisited (Squamata, Leptotyphlopidae).
–Zootaxa: 3027, pp. 63-64 - Lehr, E., Wallach, V., Köhler, G. & Aguilar, C., 2002: New species of tricolor Leptotyphlops (Reptilia: Squamata: Leptotyphlopidae) from central Peru.
–Copeia: Vol. 102, #1, pp. 131-136 - Uetz, P., Etzold, T. & Chenna, R., 1995-2015: The Reptile Database
–iNet: http://www.reptile-database.org/