After Johnson & Elliott, 1995
<==o †Coccosteomorphi Stensiö, 1944 sensu Carr & Hlavin, 2010 [Coccosteoidea sensu Johnson & Elliott, 1995] |?-o †Pholidosteidae | |-- †Malerosteus U. Dev. EEu. | |--o †Pholidosteus U. Dev. Eu. | | `-- †P. friedelti | `-- †Tapinosteus U. Dev. Eu. |--o †Coccosteidae Traquair, 1888 | |?- †Belgiosteus M. Dev. Eu. | |?-o †Clarkeosteus Obruchev, 1964; M. Dev. NA. ?U. Dev. Gr. | | `-- †C. cf. C. halmodeus (Clarke, 1893) | |?- †Jiuchengia Wang & Wang, 1983 | |-- †Livosteus M.-U. Dev. EEu. | |-- †Protitanicbthys M. Dev. NA. | |-- †Woodwardosteus [Liognathus, Lispognathus] M. Dev. NA. | |--+--o †Coccosteus M. ?U. Dev. Eu. EEu. ?NA. | | | |-- †C. decipiens | | | `-- †C. cuspidatus | | `--o †Millerosteus M. Dev. Eu. EEu. | | `-- †M. minor | `--+-- †Dickosteus M. Dev. Eu. | `--o †Watsonosteus M. Dev. Eu. | `-- †W. fletti `--+--o †Plourdostenidae Vézina, 1990 sensu Gardiner & Miles, 1994; Long, 1995 | |?- †Compagopiscis Gardiner & Miles, 1994 | |?- †Gogopiscis Gardiner & Miles, 1994 | |?-o †Plourdostenidae Vézina, 1990 sensu Long, 1995 [possibly a Panxiosteid Pachysteomorphans] | | |-- †Kimberleyichthys heintzi (Dennis-Bryan & Miles, 1983a) Dennis-Bryan & Miles, 1983b [Kimberia Dennis-Bryan & Miles, 1983a)] | | `--o †Plourdosteus Ørvig, 1951 [Pelycophorus, Tomaiosteus] U. Dev. Eu. NA. NAs. | | |-- †P. canadensis (Woodward, 1892) | | `-- †P. livonicus (Eastman) | `--o †Torosteidae Gardiner & Miles, 1990 | |?-o †Eldenosteus Miles, 1964; U. Dev. NA. | | `-- †E. arizonensis (Hussakof, 1942) [Coccosteus arizonensis Hussakof, 1942] | |--o †Harrytoombsia U. Dev. Aus. | | `-- †H. elegans | `-- †Torosteus Gardiner & Miles, 1990 `--+?-o †Incisoscutidae Denison, 1984 | `-- †Incisoscutum ritchei U. Dev. Aus. `--o †Camuropiscidae Traquair, 1888 |?- †Simosteus U. Dev. Aus. |-- †Camuropiscis U. Dev. Aus. `--+-- †Rolfosteus U. Dev. Aus. `-- †Tubonasus U. Dev. Aus.
Reference(s):
- Blom, H., Clack, J. A., Ahlberg, P. E. & Friedman, M., 2007: Devonian vertebrates from East Greenland: a review of faunal composition and distribution.
–Geodiversitas: Vol. 29, #1, pp. 119-141 - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Johnson, H. G. & Elliott, D. K., 1995: A redescription of Endenosteus arizonensis (Placodermi: Arthrodira) from the Upper Devonian Martin Formation of northern Arizona.
–Journal of Vertebrate Paleontology: Vol. 15, #2, pp. 221-234