After Holtz, 2000, and Carr, Williamson & Schwimmer, 2005
<==o †Tyrannosauroidea Osborn, 1906 sensu Holtz, 2000 [=Deinodontoidea] |-- †Guanlong wucaii Xu, Clark, Forster, Norell, Erickson, Eberth, Jia & Zhao, 2006 `--+-- †Dilong paradoxus Xu, Norell, Kuang, Wang, Zhao & Jia, 2004 `--+-- †Eotyrannus lengi Hutt, Naish, Martill, Barker & Newbery, 2001 `--+?- †Siamotyrannus isanensis Buffetaut, Suteethorn, & Tong, 1996 |?- †Labocania anomola Molnar, 1974 |--+?- †Stokesosaurus clevelandi Madsen, 1974 | `?- †Itemirus medullaris Kurzanov, 1976a `--+?- †Dryptosaurus aquilungis (Cope, 1866) [Lealaps aquilungis Cope, 1866] `--+?- †Appalachiosaurus montgomeriensis Carr, Williamson & Schwimmer, 2005 `--o †Tyrannosauridae Osborn, 1905 [=? Deinodontidae Leidy, 1856] |?- †Tyrannosauridae genera et species indet. from New Mexico [possibly abelisaurid] |--o †Aublysodontinae [probably presents juveline forms of the larger tyrannosaurines] | |?- †Aublysodontinae genera et species indet. [Kirtland Shale aublysodontine] | |-- †Aublysodon mirandis Leidy, 1868b | |-- †Alectrosaurus olseni Gilmore, 1933a | `?-o †Stygivenator [Aublysodon] | |-- †S. amplus (Marsh, 1892d) [Aublysodon amplus Marsh, 1892d] | |-- †S. cristatus (Marsh, 1892d) [Aublysodon cristatus Marsh, 1892d] | `-- †S. molnari [Aublysodon molnari] `--o †Tyrannosaurinae Osborn, 1905 [=? Deinodontinae Leidy, 1856] |?- †Tyrannosaurinae genera et species indet. [Two Medicine tyrannosaurine] |-- †Alioramus remotus Kurzanov, 1976b `--+?- †Shanshanosaurus huoyanshensis Dong, 1977 `--+-- †Albertosaurus sarcophagus Osborn, 1905 [?Deinodon, Leidy, 1856, Albertosaurus arctunguis Parks, 1928] |-- †Gorgosaurus libratus Lambe, 1914c [Albertosaurus libratus Lambe, 1914c, Lealaps incrassatus Cope, 1876a, ?Deinodon horridus Leidy, 1856, ?Lealaps faculus Cope, 1876a, ?Lealaps hazenianus Cope, 1876b, ?Dryptosaurus kenabekides Hay, 1899, ?Gorgosaurus sternbergi Matthew & Brown, 1923, a juveline form] `--o ?†Tyrannosaurini Osborn, 1905 |-- †Daspletosaurus torosus Russell, 1970a (“härkätyrannisaurus”) `--+-- †Tarbosaurus bataar (Maleev, 1955b) [taxonomic history] (“mongoliantyrannosaurus”) `-- †Tyrannosaurus rex Osborn, 1905 [taxonomic history] (“kuningastyrannisaurus”)
Reference(s):
- Buffetaut, E., Suteethorn, V. & Tong, H., 1996: The earliest known tyrannosaur from the Lower Cretaceous of Thailand.
–Nature: Vol. 381, 20 June, pp. 689-691 - Carr, T. D., Williamson, T. E. & Schwimmer, D. R., 2005: A new genus and species of tyrannosauroid from the Late Cretaceous (Middle Campanian) Demopolis Formation of Alabama.
–Journal of Vertebrate Paleontology: Vol. 25, #1, pp. 119-143 - Currie, P. J. & Dong, Z.-m., 2001: New information on Shanshanosaurus huoyanshanensis, a juveline tyrannosaurid (Theropoda, Dinosauria) from the Late Cretaceous of China.
–Canadian Journal of Earth Sciencies: Vol. 38, #12, pp. 1729-1737 - Holtz, T. R. Jr., 2000: †Tyrannosauroidea -
Tyrant dinosaurs and their possible relatives.
–iNet: The Tree of Life - Holtz, T. R. Jr., 2000: †Tyrannosauridae - Tyrant dinosaurs.
–iNet: The Tree of Life - Hutt, S., Naish, W., Martill, D. M., Barker, M. J. & Newbery, P., 2001: A preliminary account of a new tyrannosauroid theropod from the Wessex Formation (Early Cretaceous) of southern England.
–Cretaceous Research: Vol. 22, #1, pp. 227-242 - Keesey, T. M., 1996-2000: The Dinosauricon
–Internet: http://dinosauricon.com/ - Molnar, R., E., 1990: Problematic Theropoda: "Carnosaurs". 306-317
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Molnar, R. E., Kurzanov, S. M., & Dong, Z., 1990: Carnosauria. 169-209
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Paul, G. S. 1988: The Predatory Dinosaurs of the World.
–Simon and Schuster Co., New York, 1988. 464. - Xu, X., Clark, J. M., Forster, C. A., Norell, M. A., Erickson, G. M., Eberth, D. A., Jia, C. & Zhao, Q., 2006: A basal tyrannosauroid dinosaur from the Late Jurassic of China.
–Nature: Vol. 439, pp. 715-718 - Xu, X., Norell, M. A., Kuang, X., Wang, X.-l., Zhao, Q. & Jia, C., 2004: Basal tyrannosauroids from China and evidence for protofeathers in tyrannosauroids.
–Nature: Vol. 431, pp. 680-684