Mikko's Phylogeny Archive

†Neoceratosauria – advanced ceratosaurs, incl. abelisaurs

After Coria & Salgado, 1998, Padian, Hutchinson & Holtz, 1999, Coria, Chiappe & Dingus 2002, Lamanna, Martínez & Smith, 2002, Carrano, Sampson & Forster, 2002, Sereno & Brusatte, 2008, and Carrano & Sampson, 2008

<==o †Neoceratosauria Novas, 1989 [Ceratosauria Marsh, 1884a sensu stricto] (neoceratosaurs; uuskeratosaurit)
   |?- †Dandakosaurus indicus Yadagiri, 1982; E.Jur. Ind.
   |?- †“Edaphrosaurus” philtippetorum Pickering, 1995
   |?- †Chuandongocoelurus primitivus He, 1984; M.Jur. EAs.
   |?- †Genyodectes serus Woodward, 1901; E.Cret. SSAm.
   |?- †Camarillasaurus cirugedae Sánchez-Hernández & Benton, 2014; E.Cret. SWEu.
   |-- †Sarcosaurus woodi Andrews, 1921; E.Jur. WNAm.
   `--+--o †Ceratosaurus Marsh, 1884a; L.Jur. WNAm. [Ceratosauridae Marsh, 1884a(?)]
      |  |?- †C.? ingens
      |  |?- †C. dentisulcatus Madsen & Wells, 2000
      |  |?- †C. magnicornis Madsen & Wells, 2000
      |  `-- †C. nasicornis Marsh, 1884a [1888]
      `--+?-+-- †Elaphrosaurus bambergi Janensch, 1920; L.Jur. EAf.
         |  `-- †Limusaurus inextricabilis Xu, et al., 2009
         |-- †Spinostropheus gauthieri (Lapparent, 1960) Sereno, Wilson & Conrad, 2004 [Elaphrosaurus gauthieri Lapparent, 1960] E.Cret. WAf.
         `--o †Abelisauria (Bonaparte & Novas, 1985) Novas, 1992 (abelisaurs; abelisaurit)
            |?- †Ligabueino andesi Bonaparte, 1995
            `--o †Abelisauroidea (Bonaparte & Novas, 1985) Novas, 1989 (abelisauroids; abelisauruksensukuiset)
               |-- †Austrocheirus isasii Ezcurra, Agnolín & Novas, 2010; L.Cret. SSAm.
               |-- †Berberosaurus liassicus Allain, Tykoski, Aquesbi, Jalil, Monbaron, Russell & Taquet, 2007
               |?- †Compsosuchus solus Huene, 1932
               |-- †Genusaurus sisteronis Accarie, Beaudoin, Dejax, Fries, Michard & Taquet, 1995; E.Cret. WEu.
               |-- †Laevisuchus indicus Huene & Matley, 1933; L.Cret. Ind.. [paraphyletic Noasauridae??]
               |--+-- †Deltadromeus agilis Sereno, Duthiel, Iarochene, Larsson, Lyon, Magwene, Sidor, Varicchio & Wilson, 1996; L.Cret. NWAf.
               |  `--+-- †Noasaurus leali Bonaparte & Powell, 1980; L.Cret. SSAm. [Noasauridae Bonaparte & Powell, 1980]
               |     `--o †Velocisauridae Bonaparte, 1991
               |        |-- †Masiakasaurus knopfleri Sampson, Carrano & Forster, 2001; L.Cret. Mad.
               |        |?- †Santanaraptor placidus Kellner, 1999; E.Cret. NESAm. [recently treated as basal tyrannosauroid (Porfiri et al., 2014)]
               |        `-- †Velocisaurus unicus Bonaparte, 1991; L.Cret. SSAm.
               `--+?- †Ilokelesia aguadagrandensis Coria & Salgado, 1998; L.Cret. SSAm.
                  `--o †Abelisauridae Bonaparte & Novas, 1985 (abelisaurids; abelisauridit)
                     |?- †Tarascosaurus salluvicus Le Loeuff & Buffetaut, 1990
                     |?- †Pycnomenosaurus nevesi Kellner & Campos, 2002
                     |-- †Kryptops palaios Sereno & Brusatte, 2008; L.Cret. NAf.
                     `--+-- †Rugops primus Sereno, Wilson & Conrad, 2004; L.Cret. WAf.
                        `--+?- †Xenotarsosaurus bonabartei Martinez, Gimenez, Rodrigues & Bochatey, 1987
                           |?- †Abelisaurus comahuensis Bonaparte & Novas, 1985; L.Cret. SSAm.
                           `--o †Carnotaurinae
                              |-- †Ekrixinatosaurus novasi Calvo, Rubilar-Roger & Moreno, 2004; L.Cret. SSAm.
                              |-- †Indosuchus raptorius Huene & Matlay, 1933; L.Cret. Ind.
                              |--+-- †Majungasaurus crenatissimus (Deperet, 1896) [Megalosaurus crenatissimus Deperet, 1896; Majungatholus atopus Sues & Taquet, 1979] L.Cret. Mad.
                              |  |-- †Rajasaurus narmadensis Wilson et al., 2003; L.Cret. Ind.
                              |  `--o †Indosaurus Huene & Matlay, 1933; L.Cret. Ind.
                              |     |?- †I. madhyamensis
                              |     `-- †I. matleyi Huene & Matlay, 1933
                              |-- †Carnotaurinae genera et species indet. [UNPSJB-PV247] [L.Cret. SSAm.]
                              |-- †Carnotaurinae genera et species indet. [AMNH 1955] [L.Cret. Ind.]
                              `--o Carnotaurini Coria, Chiappe & Dingus 2002
                                 |-- †Aucasaurus garridoi Coria, Chiappe & Dingus, 2002; L.Cret. SSAm.
                                 `-- †Carnotaurus sastrei Bonaparte, 1985; L.Cret. SSAm.

<==o Neoceratosauria
   |-- †Limusaurus
   `--+-- †Camarillasaurus
      `--+--+-- †Deltadromeus
         |  |-- †Spinostropheus
         |  `-- †Elaphrosaurus
         `--+-- †Ceratosaurus
            `--o †Abelisauroidea
               |--o †Noasauridae
               |  |-- †Noasaurus
               |  |-- †Masiakasaurus
               |  |-- †Laevisuchus
               |  |-- †Velocisaurus
               |  `-- †Genusaurus
               `--o †Abelisauridae
                  |-- †Rugops
                  `--+-- †Ekrixinatosaurus
                     `--+--+-- †Aucasaurus
                        |  |-- †Ilokelesia
                        |  `-- †Carnotaurus
                        `--+-- †Abelisaurus
                           |-- †Indosaurus
                           |-- †Majungasaurus
                           `-- †Rajasaurus

<==o Neoceratosauria
   |-- †Deltadromeus
   |-- †Spinostropheus
   |-- †Elaphrosaurus
   `--+-- †Ceratosaurus
      `--o †Abelisauroidea
         |--o †Noasauridae
         |  |-- †Noasaurus
         |  |-- †Masiakasaurus
         |  |-- †Laevisuchus
         |  |-- †Velocisaurus
         |  `-- †Genusaurus
         `--o †Abelisauridae
            |-- †Rugops
            `--+-- †Ekrixinatosaurus
               `--+-- †Aucasaurus
                  |-- †Abelisaurus
                  |-- †Ilokelesia
                  |-- †Carnotaurus
                  `--+-- †Indosaurus
                     |-- †Majungasaurus
                     `-- †Rajasaurus

<==o Ceratosauria
   |-- †Coelophysoidea
   `--o †Neoceratosauria
      |-- †Elaphrosaurus
      `--+-- †Ceratosaurus
         `--o †Abelisauroidea
            |--o †Noasauridae
            |  |-- †Noasaurus
            |  `-- †Masiakasaurus
            `--o †Abelisauridae
               |-- †Ilokelesia
               `--+-- †Abelisaurus
                  `--+-- †Carnotaurus
                     `-- †Majungasaurus

<==o Neoceratosauria
   |-- †Elaphrosaurus
   `--+-- †Ceratosaurus
      `--+-- †Spinostropheus
         `--o †Abelisauroidea
            |?- †Genusaurus
            |?- †Laevisuchus
            |--o †Noasauridae
            |  |-- †Deltadromeus
            |  |-- Gadoufaoua noasaurid
            |  |-- †Noasaurus
            |  `-- †Masiakasaurus
            `--o †Abelisauridae
               |?- †Ilokelesia
               |-- †Rugops
               `--+-- †Abelisaurus
                  `--+-- †Rajasaurus
                     `--+-- †Majungatholus
                        `-- †Carnotaurus

Reference(s):

• Allain, R., Tykoski, R., Aquesbi, N., Jalil, N.-E., Monbaron, M., Russell, D. & Taquet, P., 2007: An abelisauroid (Dinosauria: Theropoda) from the Early Jurassic of the High Atlas Mountains, Morocco, and the radiation of ceratosaurs.
  –Journal of Vertebrate Paleontology: Vol. 27, #3, pp. 610-624 [doi: 10.1671/0272-4634(2007)27[610:aadtft]2.0.co;2]
• Benton, M. J., 1990: Origin and interrelationships of Dinosaurs. 11-30
  in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
  –University of California Press, Berkley, Los Angeles, Oxford, 1990, xvi-733
• Carrano, M. T., Sampson, S. D. & Forster, C. A., 2002: The osteology of Masiakasaurus knopfleri, a small abelisauroid (Dinosauria: Theropoda) from the Late Cretaceous of Madagascar.
  –Journal of Vertebrate Paleontology: Vol. 22, #3, pp. 510-534
• Carrano, M. T. & Sampson, S. D., 2008: The Phylogeny of Ceratosauria (Dinosauria: Theropoda).
  –Journal of Systematic Palaeontology: Vol. 6, #2, pp. (183-236) [doi: 10.1017/s1477201907002246]
• Calvo, J. O., Rubilar-Roger, D. & Moreno, K., 2004: A new Abelisauridae (Dinosauria: Theropoda) from northwest Patagonia.
  –Ameghiniana: vol. 41, #4, pp. 555-563 [from abstract in DinoData.net]
• Coria, R. A., Chiappe, L. M. & Dingus, L., 2002: A new close relative of Carnotaurus sastrei Bonaparte 1985 (Theropoda: Abelisauridae) from the Late Cretaceous of Patagonia. [= Aucasaurus]
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• Coria, R. A. & Salgado, L., 1998: A basal Abelisauria Novas, 1992 (Theropoda-Ceratosauria) from the Cretaceous of Patagonia, Argentina.
  –Gaia: nr. 15, pp. 89-102
• Ezcurra, M., Agnolin, F. L. & Novas, F. E., 2010: An abelisauroid dinosaur with a non-atrophied manus from the Late Cretaceous Pari Aike Formation of southern Patagonia.
  –Zootaxa: #2450, pp. 1-25
• Keesey, T. M., 1996-2000: The Dinosauricon
  –iNet: http://dinosauricon.com/
• Lamanna, M. C., Martínez, R. D. & Smith, J. B., 2002: A definitive abelisaurid theropod dinosaur from the Early Late Cretaceous of Patagonia.
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• Molnar, R., E., 1990: Problematic Theropoda: "Carnosaurs". 306-317
  in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
  –University of California Press, Berkley, Los Angeles, Oxford, 1990, xvi-733
• Norman, D., B., 1990: Problematic Theropoda: "Coelurosaurs". 280-305
  in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
  –University of California Press, Berkley, Los Angeles, Oxford, 1990, xvi-733
• Novas, F. E. 1996: Dinosaur monophyly.
  –Journal of Vertebrate Paleontology: vol. 16, #4, pp. 723-741.
• Padian, K., Hutchinson, J. R. & Holtz, T. R. Jr., 1999: Phylogenetic defenitions and nomenclature of the major taxonomic categories of the carnivorous Dinosauria (Theropoda).
  –Journal of Vertebrate Paleontology: Vol. 19, #1, pp. 69-80
• Paul, G. S. 1988: The Predatory Dinosaurs of the World.
  –Simon and Schuster Co., New York, 1988. 464.
• Rauhut, O. W. M., 2004: Provenance and anatomy of Genyodectes serus, a large-toothed ceratosaur (Dinosauria: Theropoda) from Patagonia.
  –Journal of Vertebrate Paleontology: Vol. 24, #4, pp. 894-902
• Rowe, T., & Gauthier, J., 1990: Ceratosauria. 151-168
  in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
  –University of California Press, Berkley, Los Angeles, Oxford, 1990, xvi-733
• Sampson, S. D., Carrano M. T. & Forster, C. A., 2000: A theropod dinosaur with bizarre dentition from the Late Cretaceous of Madagascar. [= Masiakasaurus]
  –Journal of Vertebrate Paleontology: Vol. 20, #3, suppl. to #3, 66A
• Sampson, S. D., Carrano M. T. & Forster, C. A., 2001: A bizarre predatory dinosaur from the Late Cretaceous of Madagascar.
  –Nature: Vol. 409, 25 January, pp. 504-506
• Sánchez-Hernández, B. & Benton, M. J., 2014: Filling the Ceratosaur Gap: A New Ceratosaurian Theropod from the Early Cretaceous of Spain.
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• Sereno, P. C. & Brusatte, S. L., 2008: Basal abelisaurid and carcharodontosaurid theropods from the lower Cretaceous Elrhaz Formation of Niger.
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• Xu, X., Clark, J. M., Mo, J., Choiniere, J., Forster, C. A., Erickson, G. M., Hone, D. W. E., Sullivan, C., Eberth, D. A., Nesbitt, S. J., Zhao, Q., Hernandez, R., Jia, C., Han, F.-l. & Guo, Y., 2009: A Jurassic ceratosaur from China helps clarify avian digital homologies.
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