After Carrano, Benson & Sampson, 2012
<==o †Allosauroidea (Marsh, 1878) Currie & Zhao, 1994 [Carnosauria Huene, 1920 (in partim); Allosauria Marsh, 1877b sensu Paul, 1988] (carnosaurs; karnosaurit) |--o Metriacanthosauridae Paul, 1988 [Sinraptoridae Currie & Zhao, 1993] | |-- †Xuanhanosaurus qilixiaensis Dong, 1984 | |-- †Shidaisaurus jinae Wu et al., 2009 | |--o †Yangchuanosaurus Dong, Chang, Li & Zhou, 1978 | | |?- †Y. magnus Dong, Zhou & Zhang, 1983 | | |-- †Y. shangyouensis Dong, Chang, Li & Zhou, 1978 | | `-- †Y. zigongensis (Gao, 1993) Carrano, Benson & Sampson, 2012 [Szechuanosaurus zigongensis Gao, 1993] | `--o †Metriacanthosaurinae Paul, 1988 | |-- †Metriacanthosaurus walkeri (Huene, 1923) Walker, 1964 | |-- †Siamotyrannus isanensis Buffetaut, Suteethorn, & Tong, 1996; E.Cret. SEAs. [originally treated as a tyrannosauroid (Buffetaut et al., 1996)] | `--o †Sinraptor Currie & Zhao, 1993; L.Jur. EAs. | |-- †S. dongi Currie & Zhao, 1993 | `-- †S. hepingensis (Gao, 1992) [Yangchuanosaurus hepingensis Gao, 1992] `--o Allosauria Paul, 1988 |--o †Allosauridae Marsh, 1877b [Labrosauridae marsh, 1882; Antrodemidae Stromer, 1934] | |-- †Saurophaganax maximus (Ray, 1941) Chure, 1995 [Saurophagus maximus Ray, 1941; Allosaurus amplexus (Cope, 1878b); Epanterias amplexus Cope, 1878b; Allosaurus maximus (Ray, 1941) Smith, 1998] | `--o †Allosaurus Marsh, 1877b [taxonomic history] (allosaurit, puukkohammasliskot) | |?- ?†A. sp. [Australia, so called “Polar allosaur”] | |?- †A. atrox (Marsh, 1878a) sensu Paul, 1988 (leijona-allosaurus, isoamerikanallosaurus) | |-- †A. europaeus Mateus, Walen & Antunes, 2006 [A. fragilis Peréz-Moreno et al., 1999; A. sp. Antunes & Mateus, 2003] (iberianallosaurus) | |-- †A. fragilis Marsh, 1877b (siroamerikanallosaurus, puukkohammaslisko) | `-- †a. jimmadseni Chure et al., 2006 `--o †Carcharodontosauria Benson, Carrano & Brusatte, 2009 |--o †Neovenatoridae Benson, Carrano & Brusatte, 2009 | |?- †Rapator ornitholestoides Huene, 1932 [Neovenatoridae indet. Carrano et al., 2012] | |-- †Neovenator salerii Hutt, Martill & Barker, 1996 | `--+?-o †Chiliantaisaurus Hu, 1964 [basal coelurosaurian sensu Porfiri et al. (2014)] | | |?- †C. sibiricus (Riabinin, 1914) Molnar, Kurzanov & Dong, 1990 [Allosaurus sibiricus Riabinin 1914][nomen dubium, Theropoda indet. sensu Carrano et al., 2012] | | `-- †C. tashuikouensis Hu, 1964 | `?-o †Megaraptora Benson, Carrano & Brusatte, 2009 [recently treated as tyrannosauroids (Porfiri et al., 2014)] | |?- †Orkoraptor burkei Novas, Ezcurra & Lecuona, 2008; L.Cret. SAm. | |?- †Siats meekerorum Zanno & Makovicky, 2013; L.Cret. NAm. | |--+-- †Australovenator wintonensis Hocknull, White, Tischler, Cook, Calleja, Sloan & Elliott, 2009; L.Cret. Aust. | | `-- †Fukuiraptor kitadaniensis Azuma & Currie, 2000; L.Cret. EAs. | `--+-- †Aerosteon riocoloradensis Sereno et al., 2009; L.Cret. SAm. | `-- †Megaraptor namunhuaiquii Novas, 1998; L.Cret. SAm. `--o †Carcharodontosauridae Stromer, 1931 sensu Benson, Carrano & Brusatte, 2009 and Ortega, Escaso & Sanz, 2010 (carcharodontosaurids; carcharodontosaurit, hainhammassaurit) |--+-- †Eocarcharia dinops Sereno & Brusatte, 2008; L.Cret. NAf. | `-- †Sauroniops pachytholus Cau, Dalla Vecchia & Fabbri, 2013 [published online 2012-10-27] L.Cret. NAf. `--+-- †Concavenator corcovatus Ortega, Escaso & Sanz, 2010; E.Cret. SWEu. `--+-- †Acrocanthosaurus atokensis Stovall & Langston, 1950 sensu Currie & Carpenter, 2000; E.Cret. WNAm. `--+-- †Shaochilong maortuensis (Hu, 1964) Brusatte., Benson, Chure, Xu, Sullivan & Hone, 2009; L.Cret. EAs. [Chilantaisaurus maortuensis Hu, 1964] `--o †Carcharodontosaurinae (Stromer, 1931) Carrano, Benson & Sampson, 2012 [incl. Giganotosaurinae Coria & Currie, 2006] |-- †Tyrannotitan chubutensis Novas, de Valais, Vickers-Rich & Rich, 2005; L.Cret. SSAm. `--+-- †Carcharodontosaurus saharicus (Deperet & Savornin, 1927) sensu Stromer, 1931 [Megalosaurus saharicus Deperet & Savornin, 1927, Megalosaurus africanus Huene, 1956]; L.Cret. NWAf. `--o †Giganotosaurini Coria & Currie, 2006 |?- †Unquillosaurus ceibalii Powell, 1979; L.Cret. SSAm. |-- †Giganotosaurus carolinii Coria & Salgado, 1995; L.Cret. SSAm. `-- †Mapusaurus roseae Coria & Currie, 2006; L.Cret. SSAm.
Allosauroidea incertae sedis:
†Allosaurus tendagurensis Janensch, 1925 [nomen dubium, Theropoda indet. sensu Carrano et al., 2012]
†Labocania anomola Molnar, 1974
†Erectopus Huene, 1921
†E. superbus Huene, 1932 [Megalosaurus superbus Sauvage, 1882]
†E. sauvagei (Huene, 1932)
†Megalosaurus inexpectatus del Corro, 1966; L.Cret. SSAm. [Theropoda indet., possibly Carcharodontosaurid teeth sensu Carrano et al., 2012]
†Szechuanosaurus campi Young, 1942b [Szechuanosaurus yandonensis Dong, Chang, Li & Zhou, 1987; Chiekosaurus ceratosauroides Young, 1942b [in partim]] [nomen dubium, Theropoda indet. sensu Carrano et al., 2012]
†Valdoraptor oweni (Lydekker, 1889) sensu Olshevsky, 1991 [Aristosuchus oweni??]
After Benton, 1990, Molnar, Kurzanov, & Dong, 1990, Sereno et al., 1996, Kazlev, 2001–, and Sereno & Brusatte, 2008<==o †Carnosauria |-- †Cryolophosaurus |-- †Monolophosaurus `--o †Allosauroidea |?- †Labocania |?- †Erectopus |?- †Lourinhanosaurus |--o †Sinraptoridae | |-- †Leshansaurus | |-- †Sinraptor | |-- †Szechuanosaurus | `-- †Yangchuanosaurus `--+--o †Allosauridae | |?- †Valdoraptor | |?- †Acrocanthosaurus | `--+-- †Saurophaganax | `-- †Allosaurus `--o †Carcharodontosauria |--o †Neovenatoridae | |-- †Neovenator | `--+-- †Chiliantaisaurus | `--o †Megaraptora | |?- †Orkoraptor | |--+-- †Australovenator | | `-- †Fukuiraptor | `--+-- †Aerosteon | `-- †Megaraptor `--o †Carcharodontosauridae |-- †Eocarcharia `--+-- †Concavenator `--+-- †Acrocanthosaurus `--+-- †Saochilong `--+-- †Tyrannotitan `--+-- †Carcharodontosaurus `--o †Giganotosaurinae |-- †Giganotosaurus `-- †Mapusaurus |
After Cau, Dalla Vecchia & Fabbri, 2013 (published online 2012-10-27)<==o †Carnosauria |-- †Allosaurus `--+-- †Sinraptor `--o †Carcharodontosauridae |--+-- †Eocarcharia | `-- †Sauroniops `--+-- †Acrocanthosaurus `--+-- †Saochilong `-+-- †Carcharodontosaurus `-- †Giganotosaurus |
Reference(s):
- Cau, A., Dalla Vecchia, F. M. & Fabbri, M., 2013: A thick-skulled theropod (Dinosauria, Saurischia) from the Upper Cretaceous of Morocco with implications for carcharodontosaurid cranial evolution.
–Cretaceous Research: Vol. 40, #0, pp. 251-260 [doi: http://dx.doi.org/10.1016/j.cretres.2012.09.002] - Benton, M. J., 1990: Origin and interrelationships of Dinosaurs. 11-30
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Benson, R., Carrano, M. & Brusatte, S., 2010: A new clade of archaic large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic.
–Naturwissenschaften: Vol. 97, #1, pp. 71-78 [doi: 10.1007/s00114-009-0614-x] - Brusatte, S. L., Benson, R. B. J., Chure, D. C., Xu, X., Sullivan, C. & Hone, D. W. E., 2009: The first definitive carcharodontosaurid (Dinosauria: Theropoda) from Asia and the delayed ascent of tyrannosaurids.
–Naturwissenschaften: Vol. 96, pp. 1051-1058 [doi: 10.1007/s00114-009-0565-2] - Carrano, M. T., Benson, R. B. J. & Sampson, S. D., 2012: The phylogeny of Tetanurae (Dinosauria: Theropoda).
–Journal of Systematic Palaeontology: Vol. 10, #2, pp. 211-300 [doi: 10.1080/14772019.2011.630927] - Coria, R. A. & Currie, P. J., 2002: The braincase of Giganotosaurus carolinii (Dinosauria: Theropoda) from the Upper Cretaceous of Argentina.
–Journal of Vertebrate Paleontology: Vol. 22, #4, pp. 802-811 - Coria, R. A. & Currie, P. J., 2006: A new carcharodontosaurid (Dinosauria, Theropoda) from the Upper Cretaceous of Argentina.
–Geodiversitas: Vol. 28, #1, pp. 71-118 - Coria, R. A. & Salgado, L., 1995: A new giant carnivorous dinosaur from the Cretaceous of Patagonia.
–Nature: Vol. 377, 12 September, pp. 224-226 - Currie, P. J. & Carpenter, K., 2000: A new specimen of Acrocanthosaurus atokensis (Theropoda, Dinosauria) from the Lower Cretaceous Antlers Formation (Lower Cretaceous, Aptian) of Oklohoma, USA.
–Geoversitas: Vol. 22, #2, pp. 207-246 - Franzosa, J. & Rowe, T. B., 2005: Cranial endocast of the Cretaceous theropod dinosaur Acrocanthosaurus atokensis.
–Journal of Vertebrate Paleontology: Vol. 25, #4, pp. 859-864 - Hocknull, S. A., White, M. A., Tischler, T. R., Cook, A. G., Calleja, N. D., Sloan, T. & Elliott, D. A., 2009: New Mid-Cretaceous (Latest Albian) Dinosaurs from Winton, Queensland, Australia.
–Plos One: Vol. 4, #7, pp. e6190 [doi: 10.1371/journal.pone.0006190] - Kazlev, M. A., 2001: Superfamily Allosauroidea.
–iNet: The Kheper Site: http://www.kheper.auz.com/gaia/biosphere/vertebrates/dinosauria/allosauria.htm - Keesey, T. M., 1996-2000: Carnosauria: “Meat reptiles” - Mighty Hunters
–iNet: The Dinosauricon: http://dinosauricon.com/taxa/carnosauria.html - Li, F., Peng, G., Ye, Y., Jiang, S. & Huang, D., 2009: A new carnosaur from the late Jurassic of Qianwei, Sichuan, China [in Chinese].
–Acta Geologica Sinica: Vol. 83, #9, pp. 1203-1213 - Mateus, O., Walen, A. & Antunes, M. T., 2006: The large theropod fauna of the Lourinhã Formation (Portugal) and its similarity to the Morrison Formation, with a description of a new species of Allosaurus. 1-7
in Foster J. R. & Lucas, S. G. R. M. (eds.), 2006: Paleontology and Geology of the Upper Jurassic Morrison Formation. –New Mexico Museum of Natural History and Science Bulletin: 36 - Molnar, R., E., 1990: Problematic Theropoda: “Carnosaurs”. 306-317
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Molnar, R. E., Kurzanov, S. M., & Dong, Z., 1990: Carnosauria. 169-209
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Novas, F. E. 1996: Dinosaur monophyly.
–Journal of Vertebrate Paleontology: vol. 16, #4, pp. 723-741. - Novas, F. E., Agnolín, F. L., Ezcurra, M. D., Porfiri, J. & Canale, J. I., 2013: Evolution of the carnivorous dinosaurs during the Cretaceous: The evidence from Patagonia.
–Cretaceous Research: Vol. 45, pp. 174-215 [doi: http://dx.doi.org/10.1016/j.cretres.2013.04.001] - Novas, F. E., de Valais, S., Vickers-Rich, P. & Rich, T. H. V., 2005: A large Cretaceous theropod from Patagonia, Argentina and the evolution of carcharodontosaurids.
†Naturwissenschaften: Vol. 92, #online advance publication, pp. 226-230 - Ortega, F., Escaso, F. & Sanz, J. L., 2010: A bizarre, humped Carcharodontosauria (Theropoda) from the Lower Cretaceous of Spain.
–Nature: Vol. 467, pp. 203-206 [doi: ] - Padian, K., Hutchinson, J. R. & Holtz, T. R. Jr., 1999: Phylogenetic defenitions and nomenclature of the major taxonomic categories of the carnivorous Dinosauria (Theropoda).
–Journal of Vertebrate Paleontology: Vol. 19, #1, pp. 69-80 - Paul, G. S. 1988: The Predatory Dinosaurs of the World.
–Simon and Schuster Co., New York, 1988. 464. - Porfiri, J. D., Novas, F. E., Calvo, J. O., Agnolín, F. L., Ezcurra, M. D. & Cerda, I. A., 2014: Juvenile specimen of Megaraptor (Dinosauria, Theropoda) sheds light about tyrannosauroid radiation.
–Cretaceous Research: Vol. 51, pp. 35-55 [doi: http://dx.doi.org/10.1016/j.cretres.2014.04.007] - Sampson, S. C., Witmer, L. M., Forster, C. A., Krause, D. W., O'Connor, P. M., Dobson, P. & Ravoavy, F., 1998: Predatory dinosaur remains from Madagascar: Implications for the Cretaceous biogeography of Gondwana.
–Science: Vol. 280, 15 May, pp. 1048-1051 - Sereno, P. C. & Brusatte, S. L., 2008: Basal abelisaurid and carcharodontosaurid theropods from the lower Cretaceous Elrhaz Formation of Niger.
–Acta Palaeontologica Polonica: Vol. 53, #1, pp. 15-46 [http://app.pan.pl/acta53/app53-015.pdf] - Sereno, P. C., Dutheil, D. B., Iarochene, M., Larsson, H. C. E.,Lyon, G. H., Magwene, P. M., Sidor, C. A., Verricchio, D. J. & Wilson, J. A., 1996: Predatory dinosaurs from the Sahara and Late Cretaceous faunal differentiation.
–Science: Vol. 272, 17 May, pp. 986-991 - Smith, D. G., 1998: A morphometric analysis of Allosaurus.
–Journal of Vertebrate Paleontology: Vol. 18, #1, pp. 126-142 - Zanno, L. E. & Makovicky, P. J., 2013: Neovenatorid theropods are apex predators in the Late Cretaceous of North America.
–Nat Commun: Vol. 4, pp. [doi: 10.1038/ncomms3827]