After Peters, 1997 [“The Pterosaur Homepage”]
<==o †Euthygnatha Peters, 1997 (“suoraleukaiset”) |-- †“Germanodactylus” rhamphastinus `--+--+?-o †Ornithodesmidae [incl. Gallodactylidae, Cyclorhamphidae, Caeradactylidae, and Istiodactylidae] | | |-- †Istiodactylus latidens (Seeley, 1901) Howsw, Milner & Martill, 2001 ["Ornithodesmus" latidens Seeley, 1901] [Ornithodesmidae; Istiodactylidae] | | |?- †Caeradactylus atrox | | `--o Cycnorhamphus [Gallodactylus] [Gallodactylidae] | | |-- †G. suevicus | | `-- †G. canjuersensis | `--+-- ?†Ornithocheiridae | `--o †Anhangueridae | |?-o †“Coloborhynchus” | | |-- “C.” clavirostris | | `-- “C.” wadleighi | |--o †Tropeognathus [Criorhynchus] | | |-- †T. mesembrinus | | |-- †T. robustus | | `-- †T. simus [Criorhynchus simus] | `--+-- †Arthurdactylus conan-doylensis | `--o †Anhanguera | |-- †A. blittersdorffi | `-- †A. santanae `--+--o Pterodactylus [Pterodactylidae] | |-- †P. antiquus | |-- †P. kochi | |-- †P. micronyx [Gnathosaurus subulatus??] | |-- †P. elegans | |-- †P. longicollum | |-- †P. grandis | |-- †P. grandipelvis | |-- †P. cerinensis | |-- †P. suprajurensis | |-- †P. manseli | |-- †P. pleydelli | |-- †P. arningi | |-- †P. maximus | `-- †P. sp. [?Germanodactylus cristatus] `--+--o †Ctenochasmatidae | |-- †Gnathosaurus subulatus | |--o †Ctenochasma | | |-- †C. roemeri | | |-- †C. gracile | | |-- †C. porocristata | | `-- †C. elegans | `--o †Pterodaustro | |-- †P. guinazui | `-- †P. sp. `--o Tanydiroptera Peters, 1997 |-- †Huanhepterus quingyangensis |-- †Doratorhynchus validus |-- †Diopecephalus `--o †Azhdarchidae Nesov, 1984 [Titanopterygiidae Padian, 1984] |-- †Titanopteryx philadelphiae |-- †Azhdarcho langicollis Nesov, 1984 |-- †Arambourgiania sp. |-- †Zhenjiangopterus sp. `--o †Quetzalcoatlus |-- †Q. northropi [male??] `-- †Q. sp. [female??]
Reference(s):
- Peters, D., 1997-2000: The phylogeny of pterosaurs.
–iNet: The Pterosaur homepage: The Phylogeny of Pterosaurs - Wellnhofer, P., 1991: The Illustrated Encyclopedia of Pterosaurs.
&8211;Salamander Books Ltd., London, 1991, pp. 192