†Azhdarchoidea

After Unwin, 2003

<==o †Azhdarchoidea Unwin, 1992 sensu Unwin, 2003 (tapejarids, azhdarchids and related pterosaurs)
   |?- †Aurorazhdarcho primordius Frey, Meyer & Tischlinger, 2011
   |?- †Radiodactylus langstoni Andres & Myers, 2012
   |?- †Sinopterus dongi Wang & Zhou, 2003
   |-- †Tupuxuara longicristatus Kellner & Campos, 1988; L. Cret. SA. [taxonomic history]
   `--o †Neoazhdarchia Unwin, 2003
      |--o †Tapejara Kellner, 1989 [Tapejaridae Kellner, 1989]
      |  |-- †T. wellnhoferi Kellner, 1989
      |  `--+-- †T. navigans Frey, Martill & Buchy, 2003
      |     `-- †T. imperator Campos & Kellner, 1997
      `--o †Azhdachidae Nesov, 1984  [Titanopterygiidae Padian, 1984]
         |?- †Azhdarchidae genera et species indet. [L. Cret. SWAs.] [Dalla Vecchia, Arduini & Kellner, 2001]
         |-- †Arambourgiania philadelphiae (Arambourg, 1959) Nesov & Borkin, 1989 [Titanopteryx philadelphiae Arambourg, 1959 non Titanopteryx Enderlein, 1935 (an insect)]; U. Cret. SWAs.
         |-- †Azhdarcho langicollis Nesov, 1984; U. Cret. CAs.
         |-- †Doratorhynchus validus Seeley, 1875; L. Cret. WEu.
         |-- †Hatzegopteryx thambema Buffetaut, Grigorescu & Csiki, 2002; U. Cret. EEu.
         |-- †Montanazdarcho minor Padian, de Ricqlès & Horner, 1995; U. Cret. WNA.
         |-- †Phosphatodraco mauritanicus Pereda Suberbiola, et al., 2003; U.Cret. NWAf.
         |--o †Quetzalcoatlus Lawson, 1975; U. Cret. SWNA.
         |  |-- †Q. northropi Lawson, 1975
         |  `-- †Q. sp.
         `-- †Zhejiangopterus linhaiensis Cai & Feng, 1994; U. Cret. EAs.

Alterntive phylogeny after Martill & Naish, 2006:

<==o †Azhdarchoidea Unwin, 1992 sensu Unwin, 2003 (tapejarids, azhdarchids and related pterosaurs)
   |-- †Tapejara navigans
   `--+-- †Sinopterus dongi
      `--+-- †Tapejara wellnhoferi
         `--+-- †Tapejara imperator
            `--o †Neoazhdarchia
               |-- †Tupuxuara
               `--o †Azhdachidae
                  |?- †Azhdarchidae gen. et sp. indet.
                  |-- †Arambourgiania
                  |-- †Azhdarcho
                  |-- †Doratorhynchus
                  |-- †Hatzegopteryx
                  |-- †Montanazdarcho
                  |-- †Phosphatodraco
                  |-- †Quetzalcoatlus
                  `-- †Zhejiangopterus

Reference(s):

Andres, B. & Myers, T. S., 2012: Lone Star Pterosaurs.
–Earth and Environmental Science Transactions of the Royal Society of Edinburgh: Vol. 103, #3-4, pp. 383-398 [doi: doi:10.1017/S1755691013000303]
Buffetaut, E., Grigorescu, D. & Csiki, Z., 2002: A new giant pterosaur with a robust skull from the latest Cretaceous of Romania.
–Naturwissenschaften: Vol. 89, #2, pp. 180-184
Buffetaut, E., Grigorescu, D. & Csiki, Z., 2003: Giant azhdarchid pterosaurs from the terminal Cretaceous of Transylvania (western Romania). 91-104.
in Buffetaut, E. & Mazin, J.-M., (eds.) 2003: Evolution and Palaeobiology of Pterosaurs.
–Geological Society of London, Special Publications 217, London, 2003, 1-347
Creisler, B. & Poling, J., 2002-: Pterosauria translation and pronunciation guide.
–iNet: www.dinosauria.com: http://www.dinosauria.com/dml/names/ptero.htm
Dalla Vecchia, F. M., Arduini, P. & Kellner, A. W. A., 2001: The first pterosaur from the Cenomanian (Late Cretaceous) Lagerstätten of Libanon.
–Cretaceous Research: Vol. 22, pp. 219-225
Frey, E., Martill, D. M. & Buchy, M.-C., 2003: A new species of tapejarid pterosaur with soft-tissue head crest. 65-72.
in Buffetaut, E. & Mazin, J.-M., (eds.) 2003: Evolution and Palaeobiology of Pterosaurs.
–Geological Society, Special Publications #217, London, 2003, 1-347
Frey, E., Meyer, C. & Tischlinger, H., 2011: The oldest azhdarchoid pterosaur from the Late Jurassic Solnhofen Limestone (Early Tithonian) of Southern Germany.
–Swiss Journal of Geosciences: Vol. 104, #1, pp. 35-55 [doi: 10.1007/s00015-011-0073-1]
Kellner, A. W. A. & Campos, D. d. A., 2002: The function of the cranial crest and jaws of a unique pterosaur from the Early Cretaceous of Brazil.
–Science: Vol. 297, pp. 389-392
Martill, D. M. & Naish, D., 2006: Cranial crest development in the azhdarchoid pterosaur Tupuxuara, with a review of the genus and tapejarid monophyly.
–Palaeontology: Vol. 49, #4, pp. 925-941
de Ricolès, A., Padian, K., Horner, J. R. & Francillon-Vieillot, H. 2000: Palaeohistology of the bones of pterosaurs (Reptilia: Archosauria): anatomy, ontogeny, and biomechanical implications.
–Zoological Journal of the Linnean Society: vol. 129, #3?, pp. 349-385.
Pereda Suberbiola, X., Bardet, N., Jouve, S., Iarochène, M., Bouya, B. & Amaghzaz, M., 2003: A new azhdarchid pterosaur from the Late Cretaceous phosphates of Marocco. 79-90.
in Buffetaut, E. & Mazin, J.-M., (eds.) 2003: Evolution and Palaeobiology of Pterosaurs.
–Geological Society of London, Special Publications 217, London, 2003, 1-347
Unwin, D. M., 2003: On the phylogeny and evolutionary history of pterosaurs. 139-190.
in Buffetaut, E. & Mazin, J.-M., (eds.) 2003: Evolution and Palaeobiology of Pterosaurs.
–Geological Society of London, Special Publications 217, London, 2003, 1-347
Wellnhofer, P., 1991: The Illustrated Encyclopedia of Pterosaurs.
–Salamander Books Ltd., London, 1991, pp. 1-192

Mikko's Phylogeny Archive

†Azhdarchoidea – tapejarids, azhdarchids and related pterosaurs

After Unwin, 2003

Alterntive phylogeny after Martill & Naish, 2006:

Reference(s):