Note:
Taxonomic and phylogenetic development of Aetosauria has been marred by a controversy considering taxonomic status of “Desmatosuchus” chamaensis.
As a common policy of the Archive, I have decided not to take sides at this point, but to wait the official solution of the problem, either by ICZN commission or SVP Ethics commitee.
After Parker, 2007
<==o †Aetosauria Marsh, 1884 (non Lydekker, 1889) [Aëtosauria Marsh, 1884] `--o †Stagonolepididae Lydekker, 1887 sensu Heckert & Lucas, 2000 |?- †Acompsosaurus wingatensis Mehl, 1915 |?- †Aetobarbakinoides brasiliensis Desojo, Ezcurra & Kischlat, 2012 |?- †Calyptosuchus wellesi Long & Ballew, 1985 |?- †Chilenosuchus forttae Casamiquela, 1980 |?- †Ebrachosaurus singularis Kuhn, 1936 |?- †Gorgetosuchus pekinensis Heckert, Schneider, Fraser & Webb, 2015 |?- †Polesinesuchus aurelioi Roberto-Da-Silva, Desojo, Cabreira, Aires, Müller, Pacheco & Dias-Da-Silva,, 2014 |?- †Sierritasuchus macalpini Parker, Stocker & Irmis, 2008 |?- †Stegomus arcuatus Marsh, 1896 |?- †Tecovasuchus chatterjeei Martz & Small, 2006 |--o †Desmatosuchinae Huene, 1942 sensu Heckert & Lucas, 2000 | |-- †Longosuchus meadei (Sawin, 1947) Hunt & Lucas, 1990 [Typothorax meadei Sawin, 1947] | `--+-- †Lucasuchus hunti Long & Murry, 1995 | `--+-- †Acaenasuchus geoffroyi Long & Murry 1995 [juveline Desmatosuchus? (Heckert & Lucas, 2002)] | `--o †Desmatosuchus Case, 1920; U.Trias. NA. [taxonomic history] | |-- †D. haploceros (Cope, 1892) | `-- †D. smalli Parker, 2005 [taxonomic history] `--o †Aetosaurinae Marsh, 1884 sensu Heckert & Lucas, 2000 |--+-- †Coahomasuchus kahleorum Heckert & Lucas, 1999 | |-- †Neoaetosauroides engaeus Bonaparte, 1969 | `--+--o †Aetosaurus O. Fraas, 1887 | | |-- †A. ferratus O. Fraas, 1887 | | `?- †A. crassicauda E. Fraas, 1907 [conspecific with A. ferratus sensu Parrish, 1994] | |-- †Aetosauroides scagliai Casamiquela, 1960 [Aetosauroides subsulcatus Zacarias, 1982; Aetosauroides inhamandensis Barberena et al., 1985] | `--o †Stagonolepis Agassiz, 1844 [taxonomic history] | |-- †S. robertsoni Agassiz, 1844 [taxonomic history] | |?- †S. wellesi (Long & Bellew, 1985) Murray & Long, 1989 [= Calyptosuchus] [taxonomic history] | `-- †S. olenkae Sulej, 2010 `--o †Typothoracisinae Parker, 2007 |--+--o †Redondasuchus Hunt & Lucas, 1991 | | |-- †R. reseri Hunt & Lucas, 1991 [type] | | `-- †R. rineharti Spielmann, Hunt, Lucas & Heckert, 2006 | `--o †Typothorax Cope, 1892; U.Trias. NA. | |-- †T. antiquum Lucas, Heckert & Hunt, 2002 | `-- †T. coccinarum Cope, 1892 [incl. Episcoposaurus horridus Cope, 1887] `--o †Paratypothoracisini Parker, 2007 |-- TTUP 545 `--+-- †“Desmatosuchus” chamaensis (Zeigler, Heckert & Lucas, 2002) [taxonomic history] `-- †Paratypothorax andressorum Long & Bellew, 1985
Alternative phylogenies of Aetosauria (from Parker, 2007):
Heckert et al., 2015 (fig. 5d)<==o †Aetosauria |-- †Aetosauroides `--+-- †Coahomasuchus `--o †Stagonolepididae |-- †Aetosaurus `--+-- †Neoaetosauroides `--+-- †Stagonolepis `--+-- †Calyptosuchus `--+-- †Redondasuchus `--+--o †Typothoracisinae | |-- †Typothorax | `--o †Paratypothoracisini | |-- †Tecovasuchus | `--+-- †Paratypothorax | `-- †Rioarribasuchus `--o †Desmatosuchinae |-- †Gorgetosuchus `--+-- †Longosuchus `--+-- †Sierritasuchus `--+-- †Acaenasuchus `--+-- †Lucasuchus `-- †Desmatosuchus |
Desojo et al., 2012<==o †Stagonolepididae |-- †Aetosauroides `--+-- †Coahomasuchus |-- †Neoaetosauroides |-- †Aetosaurus `--+-- †Stagonolepis |-- †Calyptosuchus `--+-- †Aetobarbakinoides `--+--o †Typothoracisinae | |--+-- †Typothorax | | `-- †Redondasuchus | `--o †Paratypothoracisini | |-- †Tecovasuchus | `--+-- †Paratypothorax | `-- †Rioarribasuchus `--o †Desmatosuchinae |-- †Sierritasuchus `--+-- †Longosuchus `--+-- †Lucasuchus `--+-- †Acaenasuchus `-- †Desmatosuchus |
Harris et al., 2003
<==o †Stagonolepididae
|-- †Aetosaurus
`--+--+-- †Aetosauroides
| `-- †Stagonolepis robertsoni
`--+--+-- †Stagonolepis wellesi [Calyptosuchus]
| `--+-- †Coahomasuchus
| `--+-- †Redondasuchus
| `-- †Typothorax
`--+-- †Neoaetosauroides
`--+-- †Paratypothorax
`--+-- †Longosuchus
`--+-- †Lucasuchus
`--+-- †Acaenosuchus
`-- †Desmatosuchus
|
Kischlat, 2000
<==o †Stagonolepididae
|-- †Aetosaurus
`--+--+-- †Coahomasuchus
| `--+-- †Redondasuchus
| `-- †Typothorax
`--+-- †Aetosauroides
|-- †Calyptosuchus
|-- †Stagonolepis
`--+-- †Neoaetosauroides
`--+-- †Paratypothorax
`--+-- †Longosuchus
`--+-- †Lucasuchus
`--+-- †Acaenosuchus
`-- †Desmatosuchus
|
Corrected typology of Heckert & Lucas, 1999 sensu Parker, 2007<==o †Stagonolepididae |--o †Aetosaurinae | `-- †Aetosaurus `--+--o †Stagonolepidinae | |-- †Aetosauroides | `-- †Stagonolepis `--+--+-- †Calyptosuchus | `--+-- †Coahomasuchus | `--+-- †Redondasuchus | `-- †Typothorax `--o Desmatosuchinae |-- †Neoaetosauroides `--+-- †Paratypothorax `--+-- †Longosuchus `--+-- †Lucasuchus `--+-- †Acaenosuchus `-- †Desmatosuchus |
Heckert & Lucas, 1999<==o †Stagonolepididae |--o †Aetosaurinae | `-- †Aetosaurus `--+--o †Stagonolepidinae | |-- †Coahomasuchus | `-- †Stagonolepis `--o †Desmatosuchinae |-- †Neoaetosauroides `--+--+-- †Longosuchus | `-- †Paratypothorax `--+-- †Typothorax `-- †Desmatosuchus |
Heckert et al., 1996
<==o †Stagonolepididae
|-- †Aetosaurus
`--+--+-- †Aetosauroides
| `-- †Stagonolepis
`--+-- †Neoaetosauroides
`--+-- †Longosuchus
|-- †Desmatosuchus
`--+-- †Paratypothorax
`--+-- †Rodondasuchus
`-- †Typothorax
|
Parrish, 1994
<==o †Stagonolepididae
|-- †Aetosaurus
`--+-- †Neoaetosauroides
|--+-- †Aetosauroides
| `-- †Stagonolepis
`--+-- †Longosuchus
`--+-- †Desmatosuchus
`--+-- †Typothorax
`-- †Paratypothorax
|
Reference(s):
- Heckert, A. B. & Lucas, S. G., 1999: A new aetosaur (Reptilia: Archosauria) from the Upper Triassic of Texas and the phylogeny of aetosaurs.
–Journal of Vertebrate Paleontology: Vol. 19, #1, pp. 50-68 - Heckert, A. B. & Lucas, S. G., 2002: South American occurences of the Adamanian (Late Triassic: Latest Carnian) index taxon Stagonolepis (Archosauria: Aetosauria) and their biochronological significance.
–Journal of Paleontology: Vol. 76, #5, pp. 852-863 - Heckert, A. B., Schneider, V. P., Fraser, N. C. & Webb, R. A., 2015: A new aetosaur (Archosauria, Suchia) from the Upper Triassic Pekin Formation, Deep River Basin, North Carolina, U.S.A., and its implications for early aetosaur evolution.
–Journal of Vertebrate Paleontology: Vol. 35, #1, pp. e881831 (1-13) [doi: 10.1080/02724634.2014.881831] - Lucas, S. G., Heckert, A. B. & Hunt, A. P., 2002: A new species of the aetosaur Typothorax (Archosauria: Stagonolepididae) from the upper Triassic of east-central New Mexico.
–New Mexico Museum of Natural History and Science Bulletin: Vol. 21, pp. 221-233 - Lucas, S. G., Hunt, A. P. & Spielmann, J. A., 2006: Rioarribasuchus, a new name for an aetosaur from the Upper Triassic of north-central New Mexico.
–New Mexico Museum of Natural History and Science Bulletin: Vol. 37, 581-582 - Martz, J. W. & Small, B. J., 2006: Tecovasuchus chatterjeei, a new aetosaur (Archosauria: Stagonolepididae) from the Tecovas Formation (Carnian, upper Triassic) of Texas.
–Journal of Vertebrate Paleontology: Vol. 26, #2, pp. 308-320 [doi: 10.1671/0272-4634(2006)26[308:tcanaa]2.0.co;2] - Parker, W. G., 2005: A new species of the Late Triassic aetosaur Desmatosuchus (Archosauria: Pseudosuchia).
–Comptes Rendus de l'Académie des Sciencies, Paris: Palevol: Vol. 4, pp. 327-340 - Parker, W. G., 2007: Reassessment of the aetosaur ‘Desmatosuchus’ chamaensis with a reanalysis of the phylogeny of the Aetosauria (Archosauria: Pseudosuchia).
–Journal of Systematic Palaeontology: Vol. 5, #1, pp. 41-68 [doi: 101017/s1477201906001994] - Parker, W. G., Stocker, M. R. & Irmis, R. B., 2008: A New Desmatosuchine Aetosaur (Archosauria: Suchia) from the Upper Triassic Tecovas Formation (Dockum Group) of Texas.
–Journal of Vertebrate Paleontology: Vol. 28, #3, pp. (692-701) [doi: 10.1671/0272-4634(2008)28[692:andaas]2.0.co;2] - Parrish, J. M., 1994: Cranial osteology of Longosuchus meadei and the phylogeny and distribution of the Aetosauria.
–Journal of Vertebrate Paleontology: Vol. 14, #2, pp. 196-209 - Roberto-Da-Silva, L., Desojo, J. B., Cabreira, S. F., Aires, A. S. S., Müller, R. T., Pacheco, C. P. & Dias-Da-Silva, S., 2014: A new aetosaur from the Upper Triassic of the Santa Maria Formation, southern Brazil.
–2014: Vol. 3764, #3, pp. 240 (39) [doi: 10.11646/zootaxa.3764.3.1] - Spielmann, J. A., Hunt, A. P., Lucas, S. G. & Heckert, A. B., 2006: Revision of Redondasuchus (Archosauria: Aetosauria) from the Upper Triassic Redonda Formation, New Mexico, with description of a new species.
in Harris, H. D., Lucas, S. G., Spielmann, J. A., Lockley, M. G., Milner, A. R. C. & Kirkland, J. L. (eds.), 2006: The Triassic-Jurassic Terrestrial Transition
–New Mexico Museum of Natural History and Science Bulletin: Vol. 37, pp. 583–587 - Sulej, T., 2010: The skull of an early Late Triassic aetosaur and the evolution of the stagonolepidid archosaurian reptiles.
–Zoological Journal of the Linnean Society: Vol. 158, #4, pp. 860-881 [doi: 10.1111/j.1096-3642.2009.00566.x] - Zeigler, K. E., Heckert, A. B. & Lucas, S. G., 2002: A new species of Desmatosuchus (Archosauria: Aerosauria) from the upper Triassic of the Chama Basin, north-central New Mexico.
–New Mexico Museum of Natural History and Science Bulletin: Vol. 21, pp. 215-219