After Dodson, & Currie, 1990, Dodson, 1996, Forster, 1996, Keesey, 1996-2000, Wu et al., 2007, Longrich, 2010, Sampson, Loewen, Farke, Roberts, Forster, Smith & Titus, 2010, Farke, Ryan, Barrett, Tanke, Braman, Loewen & Graham, 2011, and Sampson, Lund, Loewen, Farke & Clayton, 2013
Alternative phylogeny after Ryan & Russell, 2005, and Ryan, 2007
Alternative phylogeny after Longrich, 2010
Alternative phylogeny of Chasmosaurinae after Sampson et al., 2010
Alternative phylogeny of Centrosaurinae after Farke, Ryan, Barrett, Tanke, Braman, Loewen & Graham, 2011
Alternative phylogeny of Centrosaurinae after Fiorillo & Tykoski, 2012
Alternative phylogeny of Centrosaurinae after Ryan, Evans & Shepherd, 2012
Alternative phylogeny of Centrosaurinae after Sampson, Lund, Loewen, Farke & Clayton, 2013
<==o †Ceratopsidae Marsh, 1890a [Ceratopsidae Marsh, 1888 (1890) Agathaumidae Cope; Ceratopidae] |--o †Centrosaurinae Lambe, 1915 after Sampson, Lund, Loewen, Farke & Clayton, 2013 [Pachyrhinosaurinae] | |-- †Diabloceratops eatoni Kirkland & DeBlieux, 2010 | `--+--+-- †Avaceratops lammersi Dobson, 1986 | | `-- †Nasutoceratops titusi Sampson, Lund, Loewen, Farke & Clayton, 2013 | `--+-- †Albertaceratops nesmoi Ryan, 2006 | `--+--o †Centrosaurini | | |?- †Brachyceratops montanensis Gilmore, 1914b | | |?- †Monoclonius crassus Cope, 1876a | | `--+--+-- †Rubeosaurus ovatus (Gilmore, 1930) McDonald & Horner, 2010 [Styracosaurus ovatus Gilmore, 1930] | | | `-- †Styracosaurus albertensis Lambe, 1913 [?Monoclonius nasicornus Brown] | | `--+-- †Spinops sternbergorum Farke, Ryan, Barrett, Tanke, Braman, Loewen & Graham, 2011; U.Cret. WNAm. | | `--+-- †Coronosaurus brinkmani (Ryan & A.P. Russell, 2005) Ryan, Evans & Shepherd, 2012 [Centrosaurus brinkmani Ryan & A.P. Russell, 2005] | | `-- †Centrosaurus apertus Lambe, 1904b [incl. Monoclonius dawsoni, M. flexus, M. cutleri, C. longirostris] | `--+--+-- †Sinoceratops zhuchengensis Xu, Wang, Zhao & Li, 2010 | | `-- †Xenoceratops foremostensis Ryan, Evans & Shepherd, 2012 | `--o †Pachyrhinosaurini sensu Sampson, 1995b | |-- †Einiosaurus procurvicornis Sampson, 1995b | `--+-- †Achelousaurus horneri Sampson, 1995b | `--o †Pachyrhinosaurus Sternberg, 1950 | |-- †P. canadiensis Sternberg, 1950; L.Cret. NWNAm. | `--+-- †P. lakustai Currie, Langston & Tanke, 2008; L.Cret. NWNAm. | `-- †P. perotorum Fiorillo & Tykoski, 2012; L.Cret. NWNAm. `--o Chasmosaurinae Lambe, 1915 [Ceratopinae Marsh, 1890a, Ceratopsinae Marsh, 1890a emend. ??, Eoceratopsinae Lambe, 1915] |?- †Medusaceratops lokii Ryan, Russell & Hartman, 2010 |?- †Mercuriceratops gemini Ryan, Evans, Currie & Loewen, 2014 |?- †Tatankaceratops sacrisonorum Ott & Larson, 2010 |--+--+-- †Coahuilaceratops magnacuerna Loewen, Sampson, Lund, Farke, Aguillón-Martínez, de Leon, Rodríguez-de la Rosa, Getty & Eberth, 2010 | | `-- †Bravoceratops polyphemus Wick & Lehman, 2013 | `--+--+-- †Utahceratops gettyi Sampson, Loewen, Farke, Roberts, Forster, Smith & Titus, 2010 | | `-- †Pentaceratops sternbergii Osborn, 1923b | `--+?-+-- †Mojoceratops perifania Longrich, 2010 [Chasmosaurus russelli Sternberg, 1940b; Campbell et al., 2016] | | `-- †Agujaceratops mariscalensis (Lehman, 1989) Lucas, Sullivan & Hunt, 2006 [†Chasmosaurus mariscalensis Lehman, 1989] | `--o †Chasmosaurus Lambe, 1914a [Monoclonius] | |?- †C. canadiensis (Lambe, 1902) Lehman, 1990 [Monoclonius canadiensis Lambe, 1902; Ceratops canadensis (Lambe, 1902) Hatcher vide Stanton & Hatcher, 1905; Eoceratops canadensis (Lambe, 1902) Lambe, 1915] | |-- †C. russelli Sternberg, 1940b | `-- †C. belli (Lambe, 1914a) [Monoclonius belli Lambe, 1902] `--+-- †Coahuilaceratops magnacuerna Loewen, Sampson, Lund, Farke, Aguillón-MartÃnez, de Leon, RodrÃguez-de la Rosa, Getty & Eberth, 2010 `--+--+-- †Kosmoceratops richardsoni Sampson, Loewen, Farke, Roberts, Forster, Smith & Titus, 2010 | `-- †Vegaceratops irvinensis (Holmes, Forster, Ryan & Shepherd, 2001) Sampson, Loewen, Farke, Roberts, Forster, Smith & Titus, 2010 [Chasmosaurus irvinensis Holmes, Forster, Ryan & Shepherd, 2001] `--+--o †Anchiceratops Lambe, 1914a | |-- †A. ornatus Lambe, 1914a | `-- †A. sp. [Judith River Formation, North America] `--+-- †Arrhinoceratops brachyops Parks, 1925 `--o †Triceratopini Brown & Henderson, 2015 |?- †Ceratops hatcheri [C. montanus?] [nomen dubium??] |-- †Eotriceratops xerinsularis Wu, Brinkman, Eberth & Braman, 2007 |-- †Ojoceratops fowleri Sullivan & Lucas, 2010 |?- †Titanoceratops ouranos Longrich, 2011 |?- †Regaliceratops peterhewsi Brown & Henderson, 2015 `--+--o †Torosaurus Marsh, 1891b [taxonomic history] | |-- †T. latus Marsh, 1891b [taxonomic history] | `-- †T. utahensis (Gilmore, 1946b) [taxonomic history] `--+-- †Nedoceratops hatcheri (Lull, 1905) Ukrainsky, 2007 [Diceratops hatcheri Lull, 1905; Diceratus hatcheri (Lull, 1905) Mateus, 2008] `--o †Triceratops Marsh, 1889b sensu Forster, 1996 [taxonomic history] |?- †T. prorsus Marsh, 1890 [taxonomic history] `-- †T. horridus Marsh, 1889a [taxonomic history]
Alternative phylogenies of Ceratopsidae:
After Brown & Henderson, 2015<==o †Ceratopsidae |-- †Centrosaurinae `--o Chasmosaurinae |--+--+-- †Coahuilaceratops | | `-- †Bravoceratops | `--+--+-- †Utahceratops | | `-- †Pentaceratops | `--+-- †Agujaceratops | `--+-- †Mojoceratops | `-- †Chasmosaurus `--+--+-- †Anchiceratops | `-- †Arrhinoceratops `--o †Triceratopsini |-- †Regaliceratops |-- †Eotriceratops |-- †Ojoceratops `--+-- †Titanoceratops `--+-- †Nedoceratops `--+-- †Torosaurus `-- †Triceratops Reference(s):
|
After Sampson, Lund, Loewen, Farke & Clayton, 2013<==o †Ceratopsidae |-- †Chasmosaurinae `--o †Centrosaurinae |-- †Diabloceratops `--+--+-- †Avaceratops | `-- †Nasutoceratops `--+-- †Albertaceratops `--+--+--+-- †Rupeosaurus | | `-- †Styracosaurus | `--+-- †Spinops | `--+-- †Centrosaurus | `-- †Coronosaurus `--+--+-- †Sinoceratops | `-- †Xenoceratops `--+-- †Einiosaurus `--+-- †Achelousaurus `-- †Pachyrhinosaurus Reference:
|
After Ryan, Evans & Shepherd, 2012<==o †Ceratopsidae |-- †Chasmosaurinae `--o †Centrosaurinae |-- †Xenoceratops `--+-- †Diabloceratops `--+-- †Avaceratops `--+-- †Albertaceratops `--+--+-- †Spinops | `--+-- †Centrosaurus | `--+-- †Coronosaurus | `-- †Styracosaurus `--+-- †Sinoceratops `--+-- †Rupeosaurus `--+-- †Einiosaurus `--+-- †Achelousaurus `-- †Pachyrhinosaurus Reference:
|
After Fiorillo & Tykoski, 2012<==o †Ceratopsidae |-- †Chasmosaurinae `--o †Centrosaurinae |-- †Diabloceratops `--+-- †Albertaceratops |--+-- †Centrosaurus brinkmani | `--+-- †Centrosaurus apertus | `-- †Styracosaurus `--+--+-- †Einiosaurus | `-- †Rubeosaurus `--+-- †Achelousaurus `-- †Pachyrhinosaurus Reference:
|
After Farke, Ryan, Barrett, Tanke, Braman, Loewen & Graham, 2011<==o †Ceratopsidae |-- †Chasmosaurinae `--o †Centrosaurinae |?- †Brachyceratops |?- †Monoclonius |-- †Diabloceratops `--+-- †Albertaceratops |-- †Avaceratops `--+--o †Centrosaurini | |-- †Styracosaurus | |-- †Spinops | `-- †Centrosaurus `--o †Pachyrhinosaurini |-- †Einiosaurus `--+-- †Achelousaurus `-- †Pachyrhinosaurus Reference:
|
After Sampson, Loewen, Farke, Roberts, Forster, Smith & Titus, 2010<==o †Ceratopsidae |-- †Centrosaurinae `--o †Chasmosaurinae |-- †Chasmosaurus `--+-- †Mojoceratops `--+-- †Agujaceratops `--+--+-- †Utahceratops | `-- †Pentaceratops `--+-- †Coahuilaceratops `--+--+-- †Kosmoceratops | `-- †Vegaceratops `--+-- †Anchiceratops `--+-- †Arrhinoceratops `--+-- †Eotriceratops |-- †Ojoceratops `--+-- †Torosaurus `--+-- †Nedoceratops `-- †Triceratops Reference:
|
After Longrich, 2010<==o †Ceratopsidae |--o †Centrosaurinae | |-- †Avaceratops | `--+-- †Centrosaurus | `--+-- †Styracosaurus | |-- †Einiosaurus | `--+-- †Achelousaurus | `-- †Pachyrhinosaurus `--o †Chasmosaurinae |-- †Chasmosaurus `--+--+-- †Agujaceratops | `-- †Mojoceratops `--+-- †Pentaceratops `--+-- †Anchiceratops `--+-- †Arrhinoceratops `--+-- †Eotriceratops |-- †Triceratops `-- †Torosaurus Reference:
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After Ryan & Russell, 2005, and Ryan, 2007<==o †Ceratopsidae |--o †Centrosaurinae | |?- †Avaceratops | |?- †Brachyceratops | |?- †Monoclonius | |-- †Albertaceratops | `--+-- †Centrosaurus | `--+-- †Styracosaurus | `--o †Pachyrhinosaurini | |-- †Einiosaurus | `--+-- †Achelousaurus | `-- †Pachyrhinosaurus `--o Chasmosaurinae |?- †Ceratops |?- †Eotriceratops |-- †Triceratops `--+-- †Torosaurus `--+--+-- †Arrhinoceratops | `-- †Anchiceratops `--+-- †Pentaceratops `--+?- †Agujaceratops `-- †Chasmosaurus Reference(s):
|
Reference(s):
- Campbell, J. A., Ryan, M. J., Holmes, R. B. & Schröder-Adams, C. J., 2016: A Re-Evaluation of the Chasmosaurine Ceratopsid Genus
Chasmosaurus (Dinosauria: Ornithischia) from the Upper Cretaceous (Campanian) Dinosaur Park Formation of Western Canada.
–Plos One: Vol. 11, #1, pp. e0145805 [doi: 10.1371/journal.pone.0145805] - Brown, Caleb M. & Henderson, Donald M., 2015: A New Horned Dinosaur Reveals Convergent Evolution in Cranial Ornamentation in Ceratopsidae.
–Current Biology: Vol. 25, #12, pp. 1641-1648 [doi: 10.1016/j.cub.2015.04.041] - Dodson, P., 1996: The Horned Dinosaurs.
–Princeton University Press, Pinceton, New Jersey, pp. xiv-346 - Dodson, P., & Currie, P. J., 1990: Neoceratopsia. 593-618.
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Farke, A. A., Ryan, M. J., Barrett, P. M., Tanke, D. H., Braman, D. R., Loewen, M. A. & Graham, M. R., 2011: A new centrosaurine from the Late Cretaceous of Alberta, Canada, and the evolution of parietal ornamentation in horned dinosaurs.
–Acta Palaeontologica Polonica: Vol. 56, #4, pp. 691-702 [doi: 10.4202/app.2010.0121] - Fiorillo, A. R. & Tykoski, R. S., 2012: A New Maastrichtian Species of the Centrosaurine Ceratopsid Pachyrhinosaurus from the North Slope of Alaska.
–Acta Palaeontologica Polonica: Vol. 57, #3, pp. 561-573 [doi: 10.4202/app.2011.0033] - Forster, C. A., 1996: Species resolution in Triceratops: Cladistic and morphometric approaches.
–Journal of Vertebrate Paleontology: Vol. 16, #2, pp. 259-270 - Godfrey, S. J. & Holmes, R., 1995: Cranial morphology and systematics of Chasmosaurus (Dinosauria: Ceratopsidae) from the Upper Cretaceous of western Canada.
–Journal of Vertebrate Paleontology: Vol. 15, #4, pp. 726-742 [doi: 10.1080/02724634.1995.10011258] - Holmes, R. B., Forster, C. A., Ryan, M. J. & Shepherd, K. M., 2001: A new species of Chasmosaurus (Dinosauria: Ceratopsia) from the Dinosaur Park Formation of southern Alberta.
–Canadian Journal of Earth Sciencies: Vol. 38, #12, pp. 1423-1438 - Keesey, T. M., 1996-2000: The Dinosauricon
–iNet: The Dinosauricon: http://dinosauricon.com/ - Lehman, T. M., 1989: Chasmosaurus mariscalensis, sp. nov., a new ceratopsian dinosaur from Texas.
–Journal of Vertebrate Paleontology: Vol. 9, #2, pp. 137-162 - Longrich, N. R., 2011: Titanoceratops ouranos, a giant horned dinosaur from the late Campanian of New Mexico.
–Cretaceous Research: Vol. 32, #3, pp. 264-276 [doi: 10.1016/j.cretres.2010.12.007] - Longrich, N. R., 2010: Mojoceratops perifania, a new chasmosaurine ceratopsid from the late Campanian of western Canada.
–Journal of Paleontology: Vol. 84, #4, pp. 681-694 - Ryan, M. J., 2007: A new basal centrosaurine ceratopsid from the Oldman Formation, southeastern Alberta.
–Journal of Paleontology: Vol. 81, #2, pp. 376-396 [doi: 0022-3360/07/0081-376] - Ryan, M., Evans, D., Currie, P. & Loewen, M., 2014: A new chasmosaurine from northern Laramidia expands frill disparity in ceratopsid dinosaurs.
–Naturwissenschaften: Vol. 101, #6, pp. 505-512 [doi: 10.1007/s00114-014-1183-1] - Ryan, M. J., Evans, D. C. & Shepherd, K. M., 2012: A new ceratopsid from the Foremost Formation (middle Campanian) of Alberta.
–Canadian Journal of Earth Sciences: Vol. 49, #10, pp. 1251-1262 [doi: 10.1139/e2012-056] - Ryan, M. J. & Russell, A. P., 2005: A new centrosaurine ceratopsid from the Oldman Formation of Alberta and its implications for centrosaurine taxonomy and systematics.
–Canadian Journal of Earth Sciencies: Vol. 42, #7, pp. 1369-1387 - Sampson, S. D., 1995: Two new horned dinosaurs from the Upper Cretaceous Two Medicine Formation of Montana; with a phylogenetic analysis of the Centrosaurinae (Ornithischia: Ceratopsidae).
–Journal of Vertebrate Paleontology: Vol. 15, #4, pp. 743-760 - Sampson, S. D., Loewen, M. A., Farke, A. A., Roberts, E. M., Forster, C. A., Smith, J. A. & Titus, A. L., 2010: New horned dinosaurs from Utah provide evidence for intracontinental dinosaur endemism.
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–Journal of Paleontology: Vol. 79, #3, pp. 564-582 - Wick, S. & Lehman, T., 2013: A new ceratopsian dinosaur from the Javelina Formation (Maastrichtian) of West Texas and implications for chasmosaurine phylogeny.
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