After Dodson, & Currie, 1990, Dodson, 1996, Forster, 1996, Keesey, 1996-2000, Wu et al., 2007, Longrich, 2010, Sampson, Loewen, Farke, Roberts, Forster, Smith & Titus, 2010, Farke, Ryan, Barrett, Tanke, Braman, Loewen & Graham, 2011, and Sampson, Lund, Loewen, Farke & Clayton, 2013

Alternative phylogeny after Ryan & Russell, 2005, and Ryan, 2007

Alternative phylogeny after Longrich, 2010

Alternative phylogeny of Chasmosaurinae after Sampson et al., 2010

Alternative phylogeny of Centrosaurinae after Farke, Ryan, Barrett, Tanke, Braman, Loewen & Graham, 2011

Alternative phylogeny of Centrosaurinae after Fiorillo & Tykoski, 2012

Alternative phylogeny of Centrosaurinae after Ryan, Evans & Shepherd, 2012

Alternative phylogeny of Centrosaurinae after Sampson, Lund, Loewen, Farke & Clayton, 2013

<==o †Ceratopsidae Marsh, 1890a [Ceratopsidae Marsh, 1888 (1890) Agathaumidae Cope; Ceratopidae]
   |--o †Centrosaurinae Lambe, 1915 after Sampson, Lund, Loewen, Farke & Clayton, 2013 [Pachyrhinosaurinae]
   |  |-- †Diabloceratops eatoni Kirkland & DeBlieux, 2010
   |  `--+--+-- †Avaceratops lammersi Dobson, 1986
   |     |  `-- †Nasutoceratops titusi Sampson, Lund, Loewen, Farke & Clayton, 2013
   |     `--+-- †Albertaceratops nesmoi Ryan, 2006
   |        `--+--o †Centrosaurini
   |           |  |?- †Brachyceratops montanensis Gilmore, 1914b
   |           |  |?- †Monoclonius crassus Cope, 1876a
   |           |  `--+--+-- †Rubeosaurus ovatus (Gilmore, 1930) McDonald & Horner, 2010 [Styracosaurus ovatus Gilmore, 1930]
   |           |     |  `-- †Styracosaurus albertensis Lambe, 1913 [?Monoclonius nasicornus Brown]
   |           |     `--+-- †Spinops sternbergorum Farke, Ryan, Barrett, Tanke, Braman, Loewen & Graham, 2011; U.Cret. WNAm.
   |           |        `--+-- †Coronosaurus brinkmani (Ryan & A.P. Russell, 2005) Ryan, Evans & Shepherd, 2012 [Centrosaurus brinkmani Ryan & A.P. Russell, 2005]
   |           |           `-- †Centrosaurus apertus Lambe, 1904b [incl. Monoclonius dawsoni, M. flexus, M. cutleri, C. longirostris]
   |           `--+--+-- †Sinoceratops zhuchengensis Xu, Wang, Zhao & Li, 2010
   |              |  `-- †Xenoceratops foremostensis Ryan, Evans & Shepherd, 2012
   |              `--o †Pachyrhinosaurini sensu Sampson, 1995b
   |                 |-- †Einiosaurus procurvicornis Sampson, 1995b
   |                 `--+-- †Achelousaurus horneri Sampson, 1995b
   |                    `--o †Pachyrhinosaurus Sternberg, 1950
   |                       |-- †P. canadiensis Sternberg, 1950; L.Cret. NWNAm.
   |                       `--+-- †P. lakustai Currie, Langston & Tanke, 2008; L.Cret. NWNAm.
   |                          `-- †P. perotorum Fiorillo & Tykoski, 2012; L.Cret. NWNAm.
   `--o Chasmosaurinae Lambe, 1915 [Ceratopinae Marsh, 1890a, Ceratopsinae Marsh, 1890a emend. ??, Eoceratopsinae Lambe, 1915]
      |?- †Medusaceratops lokii Ryan, Russell & Hartman, 2010
      |?- †Mercuriceratops gemini Ryan, Evans, Currie & Loewen, 2014
      |?- †Tatankaceratops sacrisonorum Ott & Larson, 2010
      |--+--+-- †Coahuilaceratops magnacuerna Loewen, Sampson, Lund, Farke, Aguillón-Martínez, de Leon, Rodríguez-de la Rosa, Getty & Eberth, 2010
      |  |  `-- †Bravoceratops polyphemus Wick & Lehman, 2013
      |  `--+--+-- †Utahceratops gettyi Sampson, Loewen, Farke, Roberts, Forster, Smith & Titus, 2010
      |     |  `-- †Pentaceratops sternbergii Osborn, 1923b
      |     `--+?-+-- †Mojoceratops perifania Longrich, 2010 [Chasmosaurus russelli Sternberg, 1940b; Campbell et al., 2016]
      |        |  `-- †Agujaceratops mariscalensis (Lehman, 1989) Lucas, Sullivan & Hunt, 2006 [†Chasmosaurus mariscalensis Lehman, 1989]
      |        `--o †Chasmosaurus Lambe, 1914a [Monoclonius]
      |           |?- †C. canadiensis (Lambe, 1902) Lehman, 1990 [Monoclonius canadiensis Lambe, 1902; Ceratops canadensis (Lambe, 1902) Hatcher vide Stanton & Hatcher, 1905; Eoceratops canadensis (Lambe, 1902) Lambe, 1915]
      |           |-- †C. russelli Sternberg, 1940b
      |           `-- †C. belli (Lambe, 1914a) [Monoclonius belli Lambe, 1902]
      `--+-- †Coahuilaceratops magnacuerna Loewen, Sampson, Lund, Farke, Aguillón-Martínez, de Leon, Rodríguez-de la Rosa, Getty & Eberth, 2010
         `--+--+-- †Kosmoceratops richardsoni Sampson, Loewen, Farke, Roberts, Forster, Smith & Titus, 2010
            |  `-- †Vegaceratops irvinensis (Holmes, Forster, Ryan & Shepherd, 2001) Sampson, Loewen, Farke, Roberts, Forster, Smith & Titus, 2010 [Chasmosaurus irvinensis Holmes, Forster, Ryan & Shepherd, 2001]
            `--+--o †Anchiceratops Lambe, 1914a
               |  |-- †A. ornatus Lambe, 1914a
               |  `-- †A. sp. [Judith River Formation, North America]
               `--+-- †Arrhinoceratops brachyops Parks, 1925
                  `--o †Triceratopini Brown & Henderson, 2015
                     |?- †Ceratops hatcheri [C. montanus?] [nomen dubium??]
                     |-- †Eotriceratops xerinsularis Wu, Brinkman, Eberth & Braman, 2007
                     |-- †Ojoceratops fowleri Sullivan & Lucas, 2010
                     |?- †Titanoceratops ouranos Longrich, 2011
                     |?- †Regaliceratops peterhewsi Brown & Henderson, 2015
                     `--+--o †Torosaurus Marsh, 1891b [taxonomic history]
                        |  |-- †T. latus Marsh, 1891b [taxonomic history]
                        |  `-- †T. utahensis (Gilmore, 1946b) [taxonomic history]
                        `--+-- †Nedoceratops hatcheri (Lull, 1905) Ukrainsky, 2007 [Diceratops hatcheri Lull, 1905; Diceratus hatcheri (Lull, 1905) Mateus, 2008]
                           `--o †Triceratops Marsh, 1889b sensu Forster, 1996 [taxonomic history]
                              |?- †T. prorsus Marsh, 1890 [taxonomic history]
                              `-- †T. horridus Marsh, 1889a [taxonomic history]

Alternative phylogenies of Ceratopsidae:

After Brown & Henderson, 2015

<==o †Ceratopsidae
   |-- †Centrosaurinae
   `--o Chasmosaurinae
      |--+--+-- †Coahuilaceratops
      |  |  `-- †Bravoceratops
      |  `--+--+-- †Utahceratops
      |     |  `-- †Pentaceratops
      |     `--+-- †Agujaceratops
      |        `--+-- †Mojoceratops
      |           `-- †Chasmosaurus
      `--+--+-- †Anchiceratops
         |  `-- †Arrhinoceratops
         `--o †Triceratopsini
            |-- †Regaliceratops
            |-- †Eotriceratops
            |-- †Ojoceratops
            `--+-- †Titanoceratops
               `--+-- †Nedoceratops
                  `--+-- †Torosaurus
                     `-- †Triceratops

Reference(s):

  • Brown, C. M. & Henderson, D. M., 2015: A New Horned Dinosaur Reveals Convergent Evolution in Cranial Ornamentation in Ceratopsidae.
    –Current Biology: in press [doi: 10.1016/j.cub.2015.04.041]

After Sampson, Lund, Loewen, Farke & Clayton, 2013

<==o †Ceratopsidae
   |-- †Chasmosaurinae
   `--o †Centrosaurinae
      |-- †Diabloceratops
      `--+--+-- †Avaceratops
         |  `-- †Nasutoceratops
         `--+-- †Albertaceratops
            `--+--+--+-- †Rupeosaurus
               |  |  `-- †Styracosaurus
               |  `--+-- †Spinops
               |     `--+-- †Centrosaurus
               |        `-- †Coronosaurus
               `--+--+-- †Sinoceratops
                  |  `-- †Xenoceratops
                  `--+-- †Einiosaurus
                     `--+-- †Achelousaurus
                        `-- †Pachyrhinosaurus

Reference:

  • Sampson, S. D., Lund, E. K., Loewen, M. A., Farke, A. A. & Clayton, K. E., 2013: A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia.
    –Proceedings of the Royal Society B: Biological Sciences: Vol. 280, #1766, in press [doi: 10.1098/rspb.2013.1186]

After Ryan, Evans & Shepherd, 2012

<==o †Ceratopsidae
   |-- †Chasmosaurinae
   `--o †Centrosaurinae
      |-- †Xenoceratops
      `--+-- †Diabloceratops
         `--+-- †Avaceratops
            `--+-- †Albertaceratops
               `--+--+-- †Spinops
                  |  `--+-- †Centrosaurus
                  |     `--+-- †Coronosaurus
                  |        `-- †Styracosaurus
                  `--+-- †Sinoceratops
                     `--+-- †Rupeosaurus
                        `--+-- †Einiosaurus
                           `--+-- †Achelousaurus
                              `-- †Pachyrhinosaurus

Reference:

  • Ryan, M. J., Evans, D. C. & Shepherd, K. M., 2012: A new ceratopsid from the Foremost Formation (middle Campanian) of Alberta.
    –Canadian Journal of Earth Sciences: Vol. 49, #10, pp. 1251-1262 [doi: 10.1139/e2012-056]

After Fiorillo & Tykoski, 2012

<==o †Ceratopsidae
   |-- †Chasmosaurinae
   `--o †Centrosaurinae
      |-- †Diabloceratops
      `--+-- †Albertaceratops
         |--+-- †Centrosaurus brinkmani
         |  `--+-- †Centrosaurus apertus
         |     `-- †Styracosaurus
         `--+--+-- †Einiosaurus
            |  `-- †Rubeosaurus
            `--+-- †Achelousaurus
               `-- †Pachyrhinosaurus

Reference:

  • Fiorillo, A. R. & Tykoski, R. S., 2012: A New Maastrichtian Species of the Centrosaurine Ceratopsid Pachyrhinosaurus from the North Slope of Alaska.
    –Acta Palaeontologica Polonica: Vol. 57, #3, pp. 561-573 [doi: 10.4202/app.2011.0033]

After Farke, Ryan, Barrett, Tanke, Braman, Loewen & Graham, 2011

<==o †Ceratopsidae
   |-- †Chasmosaurinae
   `--o †Centrosaurinae
      |?- †Brachyceratops
      |?- †Monoclonius
      |-- †Diabloceratops
      `--+-- †Albertaceratops
         |-- †Avaceratops
         `--+--o †Centrosaurini
            |  |-- †Styracosaurus
            |  |-- †Spinops
            |  `-- †Centrosaurus
            `--o †Pachyrhinosaurini
               |-- †Einiosaurus
               `--+-- †Achelousaurus
                  `-- †Pachyrhinosaurus

Reference:

  • Farke, A. A., Ryan, M. J., Barrett, P. M., Tanke, D. H., Braman, D. R., Loewen, M. A. & Graham, M. R., 2011: A new centrosaurine from the Late Cretaceous of Alberta, Canada, and the evolution of parietal ornamentation in horned dinosaurs.
    –Acta Palaeontologica Polonica: Vol. 56, #4, pp. 691-702 [doi: 10.4202/app.2010.0121]

After Sampson, Loewen, Farke, Roberts, Forster, Smith & Titus, 2010

<==o †Ceratopsidae
   |-- †Centrosaurinae
   `--o †Chasmosaurinae
      |-- †Chasmosaurus
      `--+-- †Mojoceratops
         `--+-- †Agujaceratops
            `--+--+-- †Utahceratops
               |  `-- †Pentaceratops
               `--+-- †Coahuilaceratops
                  `--+--+-- †Kosmoceratops
                     |  `-- †Vegaceratops
                     `--+-- †Anchiceratops
                        `--+-- †Arrhinoceratops
                           `--+-- †Eotriceratops
                              |-- †Ojoceratops
                              `--+-- †Torosaurus
                                 `--+-- †Nedoceratops
                                    `-- †Triceratops

Reference:

  • Sampson, S. D., Loewen, M. A., Farke, A. A., Roberts, E. M., Forster, C. A., Smith, J. A. & Titus, A. L., 2010: New horned dinosaurs from Utah provide evidence for intracontinental dinosaur endemism.
    –Plos One: Vol. 5, #9, pp. e12292 (1-12) [doi: 10.1371/journal.pone.0012292]

After Longrich, 2010

<==o †Ceratopsidae
   |--o †Centrosaurinae
   |  |-- †Avaceratops
   |  `--+-- †Centrosaurus
   |     `--+-- †Styracosaurus
   |        |-- †Einiosaurus
   |        `--+-- †Achelousaurus
   |           `-- †Pachyrhinosaurus
   `--o †Chasmosaurinae
      |-- †Chasmosaurus
      `--+--+-- †Agujaceratops
         |  `-- †Mojoceratops
         `--+-- †Pentaceratops
            `--+-- †Anchiceratops
               `--+-- †Arrhinoceratops
                  `--+-- †Eotriceratops
                     |-- †Triceratops
                     `-- †Torosaurus

Reference:

  • Longrich, N. R., 2010: Mojoceratops perifania, a new chasmosaurine ceratopsid from the late Campanian of western Canada.
    –Journal of Paleontology: Vol. 84, #4, pp. 681-694

After Ryan & Russell, 2005, and Ryan, 2007

<==o †Ceratopsidae
   |--o †Centrosaurinae
   |  |?- †Avaceratops
   |  |?- †Brachyceratops
   |  |?- †Monoclonius
   |  |-- †Albertaceratops
   |  `--+-- †Centrosaurus
   |     `--+-- †Styracosaurus
   |        `--o †Pachyrhinosaurini
   |           |-- †Einiosaurus
   |           `--+-- †Achelousaurus
   |              `-- †Pachyrhinosaurus
   `--o Chasmosaurinae
      |?- †Ceratops
      |?- †Eotriceratops
      |-- †Triceratops
      `--+-- †Torosaurus
         `--+--+-- †Arrhinoceratops
            |  `-- †Anchiceratops
            `--+-- †Pentaceratops
               `--+?- †Agujaceratops
                  `-- †Chasmosaurus

Reference(s):

  • Ryan, M. J., 2007: A new basal centrosaurine ceratopsid from the Oldman Formation, southeastern Alberta.
    –Journal of Paleontology: Vol. 81, #2, pp. 376-396 [doi: 0022-3360/07/0081-376]
  • Ryan, M. J. & Russell, A. P., 2005: A new centrosaurine ceratopsid from the Oldman Formation of Alberta and its implications for centrosaurine taxonomy and systematics.
    –Canadian Journal of Earth Sciencies: Vol. 42, #7, pp. 1369-1387

 

Reference(s):