After Faivovich et al., 2005, and Frost et al., 2006, with Recent species from Frost, 2000-2002 [“AMNH Amphibian species of the World”]
<==o Lophiohylini Miranda-Ribeiro, 1926 |--o Phyllodytes Wagler, 1830 [Amphodus Peters, 1873 “1872”; Lophyohyla Miranda-Ribeiro, 1923] | |-- P. gyrinaethes Peixoto, Caramaschi & Freire, 2003 | |--o P. auratus Group | | |-- P. auratus (Boulenger, 1917) | | `-- P. wuchereri (Peters, 1873 “1872”) [Amphodus wuchereri Peters, 1873 “1872”] | |--o P. luteolus Group | | |-- P. acuminatus Bokermann, 1966 | | |-- P. brevirostris Peixoto & Cruz, 1988 | | |-- P. edelmoi Peixoto, Caramaschi & Freire, 2003 | | |-- P. kautskyi Peixoto & Cruz, 1988 | | |-- P. luteolus [Hyla luteola Wied-Neuwied, 1824; Phyllodytes luteolus (Wied-Neuwied) Wagler, 1830; Lophyohyla piperata Miranda-Ribeiro, 1923] | | `-- P. melanomystax Caramaschi, Silva & Britto-Pereira, 1992 | `--o P. tuberculosus Group | |-- P. punctatus Caramaschi & Peixoto, 2004 | `-- P. tuberculosus Bokermann, 1966 `--+-- Itapotihyla langsdorffii (Duméril & Bibron, 1841) Faivovich et al., 2005 [Hyla langsdorffii Duméril & Bibron, 1841; Osteocephalus langsdorffii (Duméril & Bibron)] `--+--+--o Osteopilus Fitzinger, 1843 [Calyptahyla Trueb & Tyler, 1974] | | |-- O. brunneus (Gosse, 1851) | | |-- O. marianae (Dunn, 1926) [Hyla marianae] | | |-- O. pulchrilineatus (Cope, 1870 “1869”) [Hyla pulchrilineata] | | |-- O. wilderi (Dunn, 1925) [Hyla wilderi] | | |--+-- O. dominicensis (Tschudi, 1838) | | | `-- O. crucialis (Harlan, 1826) [Hyla crucialis Harlan, 1826; Trachycephalus lichenatus Gosse, 1851; Calyptahyla lichenatus (Gosse) Trueb & Tyler, 1974] | | `--+-- O. septentrionalis (Duméril & Bibron, 1841) [Hyla septentrionalis Duméril & Bibron, 1841; Trachycephalus marmoratus Duméril & Bibron, 1841] | | `-- O. vastus (Cope, 1871) | `--+--o Tepuihyla Ayarzagüena & Señaris, 1993 “1992” | | |-- T. aecii (Ayarzagüena & Señaris & Gorzula, 1993 “1992”) | | |-- T. celsae Mijares-Urrutia, Manzanilla-Pupo & La Marca, 1999 | | |-- T. edelcae (Ayarzagüena & Señaris & Gorzula, 1993 “1992”) | | |-- T. galani (Ayarzagüena & Señaris & Gorzula, 1993 “1992”) | | |-- T. luteolabris (Ayarzagüena & Señaris & Gorzula, 1993 “1992”) | | |-- T. rimarum (Ayarzagüena & Señaris & Gorzula, 1993 “1992”) | | |-- T. rodriguezi (Rivero, 1968) Ayarzagüena & Señaris, 1993 [Hyla rodriguezi Rivero, 1968] | | `-- T. talbergae Duellman & Yoshpa, 1996 | `--o Osteocephalus Steindachner, 1862 | |?- O. ayarzaguenai | |-- O. buckleyi (Boulenger, 1882) | |-- O. deridens Jungfer, Ron, Seipp & Almendáriz, 2000 | |-- O. elkejungingerae (Henle, 1981) | |-- O. exophthalmus Smith & Noonan, 2001 | |-- O. fuscifacies Jungfer, Ron, Seipp & Almendáriz, 2000 | |-- O. heyeri Lynch, 2002 | |-- O. leoniae Jungfer & Lehr, 2001 | |-- O. mutabor Jungfer & Hödl, 2002 | |-- O. pearsoni (Gaige, 1929) | |-- O. planiceps Cope, 1874 | |-- O. subtilis Martins & Cardoso, 1987 | |-- O. verruciger (Werner, 1901) | |-- O. yasuni Ron & Pramuk, 1999 | |--+-- O. cabrerai (Cochran & Goin, 1970) | | `-- O. leprieurii (Duméril & Bibron, 1841) | `--+-- O. oophagus Jungfer & Schiesari, 1995 | `-- O. taurinus Steindachner, 1862 `--+--o Trachycephalus Tschudi, 1838 [Phrynohyas Fitzinger, 1843; Acrodytes Fitzinger, 1843; Scytopis Cope, 1862; Tetraprion Stejneger & Test, 1891] | |-- T. atlas Bokermann, 1966 | |-- T. coriaceus (Peters, 1867) [Phrynohyas coriacea (Peters, 1867)] | |-- I. imitatrix (Miranda-Ribeiro, 1926) [Phrynohyas imitatrix (Miranda-Ribeiro, 1926)] | |-- T. lepidus (Pombal, Haddad & Cruz, 2003) [Phrynohyas lepida Pombal, Haddad, & Cruz, 2003] | |-- T. jordani (Stejneger & Test, 1891) [Tetraprion jordani Stejneger & Test, 1891] | `--+--+-- T. mesophaeus (Hensel, 1867) [Phrynohyas mesophaea (Hensel, 1867)] | | `-- T. nigromaculatus Tschudi, 1838 | `--+-- T. hadroceps (Duellman & Hoogmoed, 1992) [Hyla hadroceps] | `--+-- T. venulosus (Laurenti, 1768) Faivovich et al., 2005 [Rana venulosa Laurenti, 1768; Hyla venulosa (Laurenti) Daudin, 1802; Acrodytes venulosa (Daudin) Fitzinger, 1843; Hyla zonata Spix, 1824] | `-- T. resinifictrix (Goeldi, 1907) [Phrynohyas resinifictrix (Goeldi, 1907)] `--+-- Corythomantis greeningi Boulenger, 1869 `--+-- Argenteohyla siemersi (Mertens, 1937) Trueb, 1970 [Hyla siemersi Mertens, 1937] `--+-- Nyctimantis rugiceps Boulenger, 1882 `--o Aparasphenodon Miranda-Ribeiro, 1920 |-- A. bokermanni Pombal, 1993 |-- A. brunoi Miranda-Ribeiro, 1920 `-- A. venezolanus (Mertens, 1950)
Reference(s):
- Faivovich, J., Haddad, C. F. B., García, P. C. A., Frost, D. R., Campbell, J. A. & Wheeler, W. C., 2005: Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic analysis.
–Bulletin of the American Museum of Natural History: #294, pp. 1-240 - Frost, D. R., 2000: Amphibian species of the World: An Online Reference.
–iNet: American Museum of Natural History, Department of Herpetology: http://research.amnh.org/herpetology/amphibia/index.html