After Frost et al., 2006, with species from Frost, 2000-2002 [“AMNH Amphibian species of the World”]

<==o Brachycephalidae Günther, 1858 (kultakonnat) [taxonomic history]
   |--o Adelophryne Hoogmoed & Lecure, 1984
   |  |-- A. adiastola
   |  |-- A. baturitensis
   |  |-- A. gutturosa
   |  |-- A. maranguapensis
   |  `-- A. pachydactyla
   |-- Atopophrynus syntomopus Lynch & Ruiz-Carranza, 1982
   |--o Barycholos Heyer, 1969
   |  |-- B. pulcher
   |  `-- B. savagei
   |--o Brachycephalus Fitzinger, 1826
   |  |-- B. ephippium (kultakonna, satulasammakko)
   |  |-- B. nodoterga
   |  `-- B. pernix
   |--o Dischidodactylus Lynch, 1979
   |  |-- D. colonnelloi
   |  `-- D. duidensis
   |--o Craugastor Cope, 1862 [Eleutherodactylus (Craugastor)]
   |  `--o C. biporcatus -species-group
   |     |-- C. aphanus (Campbell, 1994) [Eleutherodactylus aphanus Campbell, 1994]
   |     |-- C. biporcatus (W. Peters, 1863) [Strabomantis biporcatus W. Peters, 1863; Eleutherodactylus maussi auctorum (Boettger, 1893); E. cornutus Gines, 1959; Eleutherodactylus biporcatus (W. Peters, 1863)]
   |     |-- C. gulosus (Cope, 1875) [Lithodytes gulosus Cope, 1875; Eleutherodactylus gulosus (Cope)]
   |     |-- C. megacephalus (Cope, 1875) [Lithodytes megacephalus Cope, 1875; Eleutherodactylus biporcatus (W. Peters, 1863); Eleutherodactylus megacephalus (Cope)]
   |     |-- C. opimus (Savage & Myers, 2002) [Eleutherodactylus biporcatus (W. Peters, 1863); Eleutherodactylus opimus Savage & Myers, 2002]
   |     `-- E. rugosus (W. Peters, 1873) [Hylodes rugosus W. Peters, 1873; Lithodytes pelviculus Cope; Lithodytes florulentus Cope;]
   |-- “Eleutherodactylus” Duméril & Bibron, 1841 [Hylodes Fitzinger, 1843; Lithodytes Fitzinger, 1843; Strabomantis W. Peters, 1863] [655 sp.!!] [paraphyletic]
   |-- “Euhyas” Fitzinger, 1843
   |--o Euparkerella Griffins, 1959
   |  |-- E. brasiliensis
   |  |-- E. cochranae
   |  |-- E. robusta
   |  `-- E. tridactyla
   |-- Geobatrachus walkeri Ruthven, 1915
   |--o Holoaden Miranda-Ribeiro, 1920
   |  |-- H. bradei
   |  `-- H. luederwaldti
   |--o Ischnocnema Reinhardt & Lütken, 1862 “1861”
   |  |-- I. quixensis
   |  |-- I. sanctaecrucis
   |  |-- I. saxatilis
   |  |-- I. simmonsi
   |  `-- I. verrucosa
   |--o “Pelorius” Hedges, 1989 [Eleutherodactylus inoptatus -group]
   |  `-- P. inoptatus [Eleutherodactylus inoptatus -group]
   |-- Phrynopus Peters, 1873
   |--o Phyllonastes Heyer, 1977
   |  |-- P. carrascoicola
   |  |-- P. heyeri
   |  |-- P. lochites
   |  |-- P. lynchi
   |  `-- P. myrmecoides
   |-- Phyzelaphryne miriamae Heyer, 1977
   `--o Syrrhopus Cope, 1878 [incl. Tomodactylus Günther, 1900] [ Eleutherodactylus marnockii -group]
      `-- S. marnockii

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