<==o †Psammosteida Traquair, 1896 [incl. Obrucheviida Tarlo, 1964] [Psammosteidae sensu lato; incl. Drepanaspidae]
   |?- †Aspidosteus heckeri Obruchev, 1941 [Aspidophorus, Obruchevia]
   |?- †Stroshiperus
   |?- †Yoglinia bergi Obruchev, 1943
   |?-o †Obrucheviida Tarlo, 1964 sensu Tarlo, 1965
   |  `--o †Obrucheviidae Tarlo, 1964
   |     |-- †Obruchevia heckeri (Obruchev, 1936) Whitley, 1940
   |     `-- †Traquairosteus pustulatus (Traquair, 1897) Tarlo, 1964
   |--o †Drepanaspididae Traquair, 1899
   |  |--o †Drepanaspis Schlüter, 1887
   |  |  |-- †D. gemuendenensis Schlüter, 1887 [Drepanaspis gemuendensis Schlüter, 1887 (lapsus calami)]
   |  |  |-- †D. sehrieli Gross, 1933
   |  |  |-- †D. lipperti Gross, 1937
   |  |  |-- †D. jaegeri Tarlo, 1964
   |  |  |-- †D. earteri (McCoy, 1851)
   |  |  `-- †D. edwardsi Tarlo, 1964
   |  `?-o †Psephaspis Ørvig, 1961
   |     |-- †P. williamsi Ørvig, 1961
   |     `-- †P. bystrowi Tarlo, 1964
   `--+--o †Guerichosteidae Tarlo, 1964
      |  |--o †Guerichosteus Tarlo, 1964
      |  |  |-- †G. kozlowskii Tarlo, 1964
      |  |  |-- †G. kotanskii Tarlo, 1964
      |  |  |-- †G. kulczyckii Tarlo, 1964
      |  |  `-- †G. lefeldi Tarlo, 1964
      |  |--o †Hariosteus Tarlo, 1964
      |  |  |-- †H. kielanae Tarlo, 1964
      |  |  `-- †H. lobanowskii Tarlo, 1964
      |  `--o †Schizosteus Obruchev, 1940 [Cheirolepis, Microlepis]
      |     |-- †S. asatkini Obruchev, 1940
      |     |-- †S. heterolepis (Preobrajensky 1911)
      |     |-- †S. torien sis Mark-Kurik, 1965
      |     |-- †S. striatus (Gross, 1933)
      |     |-- †S. perneri (Ruzicka, 1929)
      |     `-- †S. wellsi Tarlo, 1964
      `--+--+--o †Psammolepis Agassiz, 1845 [Psammolepididae Tarlo, 1962]
         |  |  |-- †P. paradoxa (Agassiz, 1845)
         |  |  |-- †P. venyukovi Obruchev, 1965
         |  |  |-- †P. alata Obruchev, 1965
         |  |  |-- †P. heteraster Gross
         |  |  |-- †P. undulata (Agassiz, 1845)
         |  |  |-- †P. proia Mark-Kurik 1965
         |  |  |-- †P. abavica Mark-Kurik, 1965
         |  |  |-- †P. aerata Obruchev, 1965
         |  |  |-- †P. arctica (Kiaer, 1915)
         |  |  |-- †P. groenlandica Tarlo, 1964
         |  |  `-- †?P. granulata (McCoy. 1848)
         |  `--o Pycnosteidae Tarlo, 1962
         |     |--o †Ganosteus Rohon, 1901
         |     |  |-- †G. stellatus Rohon, 1901
         |     |  |-- †G. artus Mark-Kurik, 1965
         |     |  `-- †G. obtusus Mark-Kurik, 1965
         |     |--o †Pycnolepis Tarlo, 1964
         |     |  `-- †P. splendens (Eichwald, 1844)
         |     |--o †Pycnosteus Preobrajensky, 1911
         |     |  |-- †P. palaeformis Preobrajensky, 1911
         |     |  |-- †P. pauli Mark, 1956
         |     |  |-- †P. tuberculatus (Rohon, 1901)
         |     |  |-- †P. nathorsti Obruchev, 1965
         |     |  `-- †P. obruchevi Tarlo, 1964		 
         |     `--o †Tartuosteus Obruchev, 1961 [=? Psammolepis]
         |        |-- †T. giganteus (Gross, 1933)
         |        |-- †T. luhai Mark-Kurik, 1965
         |        `-- †T. maximus Mark-Kurik, 1965
         `--o Psammosteidae Traquair, 1896 sensu stricto
            |-- †Crenosteus levis (Obruchev, in Tarlo, 1964) Tarlo, 1964
            |-- †Karelosteus weberi Obruchev, 1933
            |--o †Psammosteus Agassiz, 1845 [Dyptychosteus, Megalopteryx, Placosteus]
            |  |-- †P. megalopteryx (Trautschold, 1880)
            |  |-- †P. maeandrinus Agassiz, 1845
            |  |-- †P. praecursor Obruchev, 1947
            |  |-- †P. markae Tarlo, 1961
            |  |-- †P. falcatus Gross, 1942
            |  |-- †P. waltergrossi Tarlo, 1965
            |  |-- †P. pectinatus Obruchev, 1965
            |  |-- †P. asper Obruchev, 1965
            |  |-- †P. tchernovi Obruchev, 1965
            |  `-- †P. kiaeri Tarlo, 1964
            `--o †Rohonosteus Tarlo, 1964
			   |-- †R. ornatus (Rohon, 1899)
               `-- †R. complicatus (Kjaer, 1915)

Reference(s):

  • Carroll, R. L., 1988: Vertebrate paleontology and evolution.
    –W. H. Freeman and company, New York, 1988, 698
  • Carroll, R. L., 1988: Appendix. 594-648
    in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
    –W. H. Freeman and company, New York, 1988, 698
  • Elliott, D. K. 1983: New Pteraspididae (Agnatha, Heterostraci) from the Lower Devonian of Northwest Territories, Canada.
    –Journal of Vertebrate Paleontology: vol. 2, #4, pp. 389-406
  • Frickhinger, K. A., 1995: Fossil Atlas – Fishes.
    –Mergus – Publishers for Natural History and Pet Books, Hans A. Baensch, Malle, Germany, 1-1088
  • Janvier , P., 1997: Vertebrata: Animals with backbones.
    –iNet: Tree of Life: http://tolweb.org/tree/
  • Lundberg, J. G., 1995, 1997: Chordata
    –iNet: Tree of Life: http://tolweb.org/tree/
  • Long, J. A., 1995: The Rise of Fishes: 500 million years of evolution.
    –Johns Hopkins University Press, Baltimore & London, pp. 1-223
  • Nelson, J. S., 1994: Fishes of the world.
    –John Wiley & Sons Inc., New York, 1994, xx-600
  • Tarlo, L. B. H., 1965: Psammosteiformes (Agnatha) a review with descriptions of new material from the lower Devonian of Poland II. Systematic part.
    –Palaeontologica Polonica: #15, pp. IX-166 (with 48 text-figures and 19 plates) [http://palaeontologia.pan.pl/Archive/1965-15.pdf]
  • White, T., 2001: Unit 1.2: Pteraspidomorphi
    –iNet: The Vertebrate Notes: http://home.houston.rr.com/vnotes/unit1.2.html