After de Pinna, 1996, Arratia, 2000, and Arratia, 2013
<==o Teleostei Müller, 1844 [Teleosteomorpha Arratia, 2000 sensu Arratia, 2013, 2017] (modernit viuhkaeväiset) |?- †Chongichthys dentatus Arratia, 1982, U.Jur. SA. [Chongichthyidae] |--+-- †Pachycormiformes Berg, 1940 | `-- †Aspidorhynchiformes Bleeker, 1859 `--+-- †Prohalecites porroi (Bellotti, 1857) Deecke, 1889 [Pholidophorus porro Bellotti, 1857; Prohalecites porro (Bellotti) Deecke, 1889; Prohalecites porroi (Bellotti) De Alessandri, 1910] M.-L.Trias. Eu. [Prohaleciteiformes Arratia, 2017: Prohaleciteidae Arratia, 2017] `--o Teleostei sensu Arratia, 2013 `==o †“Pholidophoriformes” Berg, 1940 [paraphyletic grade of basal teleosteans] |--o †Pholidophoridae Woodward, 1890 sensu Arratia, 2013 [Pholidophoriformes Berg, 1940 sensu stricto] | |?-o †Malingichthys Tintori, Sun, Ni, Lombardo, Jiang & Motani, 2015; M.Trias. SEAs. | | |-- †M. nimaiguensis Tintori, Sun, Ni, Lombardo, Jiang & Motani, 2015 | | `-- †M. wanfenglinensis Tintori, Sun, Ni, Lombardo, Jiang & Motani, 2015 | |?- †Zambellichthys bergamensis Arratia, 2013; L.Trias. SEu. | |-- †Annaichthys pontegiurinensis Arratia, 2013; L.Trias. SEu. | `--+--+-- †Pholidophoretes salvus Griffith, 1977; L.Trias. SEu. | | `--o †Pholidoctenus Zambelli, 1977; L.Trias. SEu. | | |-- †P. serianus Zambelli, 1977; L.Trias. SEu. | | `-- †P. sanpellegrinensis Arratia, 2017; L.Trias. SEu. | `--+--o †Parapholidophorus Zambelli, 1975; L.Trias. SEu. | | |-- †P. caffi (Airaghi, 1908) [Pholidophorus caffi Airaghi, 1908] | | `-- †P. nybelini Zambelli, 1975 [type] | `--+-- †Pholidophorus latiusculus Agassiz, 1832 [P. fusiformis Agassiz, 1844] [Baleiichthys; ?Microps; ?Nothosomus; Phelidophorus; Poreirgus] ? E.Trias.-L.Jur. SAm., M.Trias.-L.Jur. Eu., L.Trias.-L.Jur. WAf., E.Jur. NAm., L.Jur. As. | `--+-- †Lombardichthys gervasuttii (Zambelli, 1980a) Arratia, 2017 [Pholidophorus gervasuttii Zambelli, 1980a] | `--+-- †Pholidorhynchodon malzanni Zambelli, 1980b [Pholidorhynchodon malzannii Arratia, 2017]; L.Trias. SEu. | `-- †Knerichthys bronni (Kner, 1866) Arratia, 2013 [Pholidophorus bronni Kner, 1866] L.Trias. SEu. `--+-- †Eurycormus speciosus Wagner, 1863 [Eurycormidae Arratia, 2017] L.Jur. Eu. `--+--o †Catervariolidae Saint-Seine, 1955 [Catervarioliformes Taverne, 2013a] | |-- †Catervariolus hornemani Saint-Seine, 1955; L.Jur. WAf. | `--+-- †Songanella callida Saint-Seine, 1955; L.Jur. WAf. | `-- †Kisanganichthys casieri Taverne, 2014; L.Jur. WAf. `--+--o †Ankylophoridae Gaudant, 1978 [Siemensichthys -group (Arratia, 2013); Ankylophoriformes Taverne, 2013b] | |?- †“Pholidophorus” germanicus Quenstedt, 1858 | |?- †Songaichthys luctacki Taverne, 2013; M.Jur. CAf. | |?- †Steurbautichthys aequatorialis (Saint-Seine & Casier, 1962) Taverne, 2011 [Pholidophorus aequatorialis Saint-Seine & Casier, 1962]; M.Jur. CAf. | |?-o †Pholidophoristion Woodward, 1941; L.Jur.-E.Cret. Eu. | | |-- †P. ornatus (Agassiz, 1844) | | |-- †P. micronyx (Agassiz, 1844) | | |-- †P. ovatus (Wagner, 1860) | | `-- †P. spaethi (Taverne, 1981) | |?- †Neopholidophoropsis serrata Taverne, 1981; E.Cret. CEu. | |?- †Lemanophorus | |-- †Lehmanophorus segusianus (Saint-Seine, 1949) Gaudant, 1978; L.Jur. WEu. | `--+-- †Ankylophorus similis (Woodward, 1895) Gaudant, 1978; L.Jur. WEu. | `--o †Siemensichthys Arratia, 2000; E.Cret. CEu. | |-- †S. macrocephalus (Agassiz, 1833-44) [Pholidophorus macrocephalus Agassiz, 1833-44] | `-- †S. siemensi Arratia, 2000 `--+-- †Dorsetichthys bechei (Agassiz, 1837) Arratia, 2013 [Pholidophorus bechei Agassiz, 1837; Dorsetichthts (sic) Arratia, 2013 (lapsus calami)] [Dorsetichthyiformes Nelson, Grande & Wilson, 2016: Dorsetichthyidae Nelson, Grande & Wilson, 2016] `--+--o †Ichthyokentemidae Griffith & Patterson, 1963 | |-- †Ichthyokentema purbeckensis (Davies, 1887) Woodward, 1941; L.Jur. Eu. [Pholidophorus purbeckensis Davies, 1887] | `?-o †Elpistoichthys Griffith, 1977; L.Trias. Eu. | |-- †E. pectinatus Griffith, 1977 | `-- †E. striolatus Griffith, 1977 `--+--o †Proleptolepis Nybelin, 1974; E.Jur. Eu. [3 sp.] | |-- †P. sp.1 | |-- †P. sp.2 | `-- †P. elongata `--+--o †Leptolepididae Pictet, 1854 [†Leptolepidiformes Nelson, 2006; “Leptolepiformes”] | |-- †Longileptolepis wiedenrothi Arratia, 2003 [Paraleptolepis Arratia & Thies, 2001 (preocc.)] | `--o †Leptolepis Agassiz, 1843 [Liassolepis; Megastoma; Sarginites] L.Jur.-L.Cret. Eu. NAm., Jur. Spits. Aust., Jur.-Cret. As. ?Jur.- E.-L. Cret. SAm. E.-L.Cret. Af. | |-- †L. africana Gardiner, 1960 | |-- †L. autissiodorensis Sauvage, 1892 | |-- †L. brodiei Agassiz, 1845 [Leptolepis nanus Egerton, 1845; Leptolepis neocomiensis Bassani, 1879; Megastoma apenninum Costa, 1850; Oxygonius tenuis Agassiz, 1845; Sarginites pygmaeus Costa, 1850] | |-- †L. concentricus Egerton, 1852 | |-- †L. coryphaenoides (Bronn, 1830) [Cyprinus coryphaenoides Bronn, 1830; Leptolepis affinis Sauvage, 1874; Leptolepis bronni Agassiz, 1833; Leptolepis constrictus Egerton, 1852; Leptolepis pachystelus Sauvage, 1874; Leptolepis pronus Sauvage, 1874] [type] | |-- †L. disjectus Woodward, 1890 | |-- †L. jaegeri Agassiz, 1844 | |-- †L. knorrii | |-- †L. lowei Woodward, 1895 | |-- †L. macrophthalmus Egerton, 1853 | |-- †L. nevadensis David, 1941 | |-- †L. normandica Nybelin, 1962 | |-- †L. opercularis Arratia, Chang & Chong, 1975c | `-- †L. saltviciensis Simpson, 1884 `--+?-o †Luisiellidae Sferco, López-Arbarello & Báez, 2015 | |-- †Waldmanichthys koonwarrensis (Waldman, 1971) Sferco, López-Arbarello & Báez, 2015 [Leptolepis koonwarrensis Waldman, 1971; Leptolepis koonwarri Waldman, 1971] | `--+-- †Cavenderichthys talbragarensis (Woodward, 1895) Arratia, 1997 | `-- †Luisiella feruglioi (Bordas, 1942) Bocchino, 1967 [incl.? Luisiella inexcutata Bocchino, 1967] `==o †Ascalaboidae Arratia, 2016 [paraphyletic sensu Arratia, 2013, monophyletic in Arratia, 2017] |-- †Ebertichthys ettlingensis Arratia, 2016; L.Jur. CEu. |-- †Ascalabos voithii Graf zu Münster, 1839b [Leptolepis polyspondylus Agassiz, 1843; Leptolepis paucispondylous Agassiz, 1843 (Nomen nudum)] L.Jur. CEu. `--+-- †Tharsis dubius (De Blainville, 1818) Giebel, 1848 [Leptolepis dubia De Blainville, 1818] L.Jur. Eu. `--+--o †Varasichthyidae Arratia, 1981 [included in Crossognathiformes by Taverne (1989).] | |-- †Domeykos profetaensis L.Jur. SA. | `--+-- †Luisichthys vinalensis White, 1942; L.Jur. WInd. | `--+-- †Varasichthys ariasi Arratia, 1981, L.Jur. SAm. | `--o †Protoclupea | |-- †P. atacamensis Arratia & Schultze, 1985 | |-- †P. chilensis Arratia, Chang & Chong | `?- †P. sp. `-- Teleocephala de Pinna, 1996 (modernit viuhkaeväiset; modern teleosteans)
Teleostei incertae sedis:
†Bunoderma baini Dolgopol de Sáez, 1940b; L.Jur. SESAm. [originally described as coelacanth]
†“Chanoides” weberi Taverne & de Cosmo, 2005 (nomen dubium) L.Cret. SEu. (Mayrinck et al., 2014)
†Clupavichthys dufouri Gayet, 1989
†Eopholidophorus forojuliensis Zambelli, 1989; L.Trias. SEu.
†Neoproscinetes De Figueiredo & Silva Santos, 1990; E.Cret. SAm. [Santana Form.]
†N. penalvai (Silva Santos, 1970)
†“Pholidophorus” vallejensis Rusconi, 1947; ?M.Trias. SAm.
†“Pholidophorus” dentatus L.Trias. SAm.
†Paraleptolepis Yabumoto, 1994;
†Paraleptolepis kikuchii Yabumoto, 1994
†Paraleptolepis elegans Yabumoto, 1994
†“Pholidophoriformes” Berg, 1940
†Hulettia americana (Eastman, 1899) Schaeffer & Patterson, 1984 [Pholidophorus americanus Eastman, 1899] L.Jur. NA.
†Lophiostomus dixoni Egerton, 1852 [a ionoscopiform?]
†Luxembourgichthys friedeni (Delsate, 1999) Taverne & Steurbaut, 2017 [Pholidophorus friedeni Delsate, 1999]
†Pholidophorichthys inermis Gaudaut, 1978; L.Jur. WEu.
†Galkiniidae Yakovlev, 1962
†Caramurus
†Galkinia Ghekker, 1948 [non Galkinia Ross & Newman, 1995 (Crustacea: Cirripedia: Pyrgomatidae)]
†Majokidae Saint-Seine, 1955;
†Majokia brasseuri Saint-Seine, 1955; L.Jur. WAf.
†Lingulellidae Saint-Seine, 1955
†Lingulella Saint-Seine, 1955; L.Jur. CAf.
†Ligulella sluysi Saint-Seine, 1955
†Ligulella fourmarieri
†Lombardinidae Saint-Seine, 1955
†Lombardina decorataSaint-Seine, 1955;
†Archaeomaenidae Goodrich, 1909 [are pachycormiforms?]
†Archaeomaene tenuis Woodward, 1895; L.Jur. Aus.
†Aetheolepis mirabilis Woodward, 1895 [Aetheolepidae Wade, 1941] L.Jur. Aus.
†Aphnelepis L.Jur. Aus. [Aphnelepidae Wade, 1941]
†Gurvanichthys mongoliensis Jakovlev, 1986; E.Jur. EAs.
†Zaxilepis qinglongensis Su, 1994 E.Jur. EAs.
†Oreochima ellioti Schaeffer, 1972 [non Schaeffer & Elliot, 1972] E.Jur. Ant.
†Madariscus robustus (Wade, 1941) [Archaeomaene robustus Wade, 1941] L.Jur. Aus.
†Wadeichthys oxyops Waldman, 1971 E.Cret. Aus.
†Koonwarria manifrons Waldman, 1971 [Koonwarriidae] E.Cret. Aust.
†Oligopleuridae Woodward, 1895
†Callopterus agassizi L.Jur. Eu.
†Oligopleurus esocinus L.Jur.-E.Cret. Eu., L.Cret. SAm.
†Spathiurus dorsalis Davis [Amphilaphurus major Davis] L.Cret. SWAs.
†Pholidophoridae Woodward, 1890
†Flugopterus Whitley, 1951 [Megalopterus Kner, 1866 (preocc.?] L.Trias. Eu.
†Hungkiichthys Liu & Wang, 1961; L.Jur. EAs.
†Jialingichthys Su, 1983 E.Jur. EAs.
†Pholidolepis dorsetensis Nybelin, 1966 E.Jur. Eu.
†Pholidophorus Agassiz, 1832 [Baleiichthys; ?Microps; ?Nothosomus; Phelidophorus; Poreirgus] ? E.Trias.-L.Jur. SAm., M.Trias.-L.Jur. Eu., L.Trias.-L.Jur. WAf., E.Jur. NAm., L.Jur. As.
†Pholidophorus australis Woodward, 1908
†Pholidophorus barazzettii Bassani, 1886
†Pholidophorus brevis Davies, 1887
†Pholidophorus curionii (Heckel, 1849) [Palaeoniscus curioni Heckel, 1849]
†Pholidophorus domeykanus Arratia, Chang & Chong, 1975a
†Pholidophorus dubius Woodward, 1890
†Pholidophorus gaudryi Sauvage, 1892
†Pholidophorus germanicus Quenstedt, 1858
†Pholidophorus granulatus Egerton, 1854
†Pholidophorus gregarius Woodward, 1890
†Pholidophorus hartmanni Egerton, 1843
†Pholidophorus higginsi Egerton, 1854
†Pholidophorus indicus
†Pholidophorus kingii
†Pholidophorus microlepidotus Kner, 1866
†Pholidophorus nitidus Egerton, 1854
†Pholidophorus pachysomus Egerton, 1852
†Pholidophorus pusillus Agassiz, 1832 [Pholidophorus knerii de Zigno, 1891] [incertae sedis, type material lost]
†Pholidophorus stricklandi Agassiz, 1844
†Pholidophorides Woodward, 1941 E.Jur. Eu.
†Pleuropholidae Saint-Seine, 1949
†Gondwanapleuropholis longimaxillaris Brito & Gallo, 2002; U.Jur. ESAm.
†Zurupleuropholis Giordano, Succar, Codorniú, Cione & Arratia, 2018; E.Cret. SSAm.
†Zurupleuropholis quijadensis Giordano, Succar, Codorniú, Cione & Arratia, 2018; E.Cret. SSAm.
†Zurupleuropholis decollavi Giordano, Succar, Codorniú, Cione & Arratia, 2018; E.Cret. SSAm.
†Austropleuropholis Saint-Seine, 1955; L.Jur. WAf.
†Parapleuropholis Saint-Seine, 1955; L.Jur. WAf.
†Pleuropholis Egerton, 1857; L.Jur. WAf., L.Jur.-E.Cret. Eu.
†Pleuropholis attenuatus (Morris, 1854) Egerton, 1858 [attenuata Morris, 1854]
†Pleuropholis crassicauda Egerton, 1858
†Pleuropholis formosa Woodward, 1919
†Pleuropholis laevissimus
†Pleuropholis longicaudus Egerton, 1858
†Pleuropholis serrata Egerton, 1858
†Signeuxellidae Saint-Seine, 1955;
†Signeuxella significans Saint-Seine, 1955;
†Siyuichthyidae Zhang & Su, 1985
†Siyuichthys Su, 1980
†Siyuichthys pulcher Su, 1980
†Siyuichthys ornatus Su, 1985
†Siyuichthys pulchellus Su, 1985
†Wukungia houyanshanensis Su, 1985
†Dsungarichthys bilineatus Su, 1985
†Bogdaichthys Su, 1985
†Bogdaichthys serraus Su, 1985
†Bogdaichthys fukangensis Su, 1985
†Manasichthys Su, 1985
†Manasichthys elongatus Su, 1985
†Manasichthys tuguluensis Su, 1985
†“Leptolepiformes”
†“Leptolepidae” Nicholson & Lydekker, 1889
†Carsothrissops D'Erasmo 1946; L.Cret. Eu.
†Cteniolepis L.Jur. Eu.
†Eurystichthys Whitley, 1950 [Eurystethus] L.Jur. Eu.
†Tharrias araripis Jordan & Branner; L.Cret. SAm.
†Todiltia schoewi Schaeffer & Patterson, 1984; M.Jur. NAm.
†Cavenderichthys talbragarensis (Woodward, 1895) Arratia, 1997 [Leptolepis talbragarensis Woodward, 1895]
†Paraclupavus caheni de Saint-Seine & Casier, 1962
After Arratia, 2017<==o Teleosteomorpha |--+-- †Pachycormiformes Berg, 1940 | `-- †Aspidorhynchiformes Bleeker, 1859 `--+--o †Prohaleciteiformes | `--o †Prohaleciteidae | `-- †Prohalecites porroi `--o Teleostei |--o †Pholidophoriformes | |--o †Eurycormidae | | `-- †Eurycormus speciosus | `--o †Pholidophoridae | |-- †Annaichthys | `--+--+-- †Pholidophoretes | | `--+-- †Malingichthys | | `--o †Pholidoctenus | | |-- †P. sanpellegrinensis | | `-- †P. serianus | `--+-- †Parapholidophorus | `--+-- †Pholidophorus latiusculus | `--+-- †Pholidorhynchodon | `-- †Lombardichthys `--+--o †Ichthyokentemidae | `-- †Ichthyokentema |--o †Ankylophoridae | |-- †Siemensichthys siemensi | `--+-- †Siemensichthys macrocephalus | `--+-- †Lehmanophorus | `-- †Ankylophorus `--+--o †Dorsetichthyiformes | `--o †Dorsetichthyidae | `-- †Dorsetichthys bechei `--+--o †Leptolepidiformes | `--o †Leptolepididae | `-- †Leptolepis coryphaenoides `--+--o †Ascalaboidae | |-- †Tharsis | `--+-- †Ebertichthys | `-- †Ascalabos `--+--o †Varasichthyidae | |-- †Domeykos | `--+-- †Varasichthys | `--+-- †Luisichthys | `-- †Protoclupea `-- Teleocephala Reference(s):
|
After Arratia, 2013<==o Teleosteomorpha |--+-- †Pachycormiformes Berg, 1940 | `-- †Aspidorhynchiformes Bleeker, 1859 `--+-- †Prohalecites porroi `--o Teleostei |--o †Pholidophoridae | |-- †Annaichthys | `--+--+-- †Pholidophoretes | | `-- †Pholidoctenus | `--+-- †Parapholidophorus | `--+-- †Pholidophorus latiusculus | `--+-- †Pholidophorus gervasuttii | `--+-- †Pholidorhynchodon | `-- †Knerichthys `--+-- †Eurycormus `--+-- †Catervariolus `--+--o Siemensichthys -group | |-- †Lehmanophorus | `--+-- †Ankylophorus | `--o †Siemensichthys | |-- †S. macrocephalus | `-- †S. siemensi `--+-- †Dorsetichthys bechei `--+-- †Ichthyokentema `--+-- †Leptolepis coryphaenoides `--+-- †Ascalabos `--+-- †Tharsis `--+--o †Varasichthyidae | |-- †Domeykos | `--+-- †Varasichthys | `--+-- †Luisichthys | `-- †Protoclupea `-- Teleocephala Reference(s):
|
After Taverne, 2011<==o Teleostei |-- †Pachycormiformes Berg, 1940 `==o “Pholidophoriformes” |-- †Catervariolus `--+--o †Ankylophoridae | |--+-- †Eurycormus | | `-- †Neopholidophoropsis | `--+-- †Steurbautichthys | `--+-- †“Pholidophorus” germanicus | `--+-- †Pholidophoristion | `--+-- †Siemensichthys | `--+-- †Ankylophorus | `-- †Lehmanophorus `--+--o †Ichthyokentemidae | `-- †Ichthyokentema `--+--o †Pholidophoridae | |-- †Pholidophorus | `?-+-- †Pholidophoretes | `-- †Pholidoctenus `--+--+-- †Archaeomaenidae | `--+-- †Aetheolepidae | `-- †Aphnelepidae `--+-- †Pleuropholidae `--+-- †“Pholidophorus” netchkini `--+-- †“Pholidophorus” domeykanus `--+-- †Pholidolepis `--+-- †Proleptolepis `--+-- †Longileptolepis `--+-- †Leptolepis `-- “Autres téléostéens” Reference(s):
|
Reference(s):
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