After Nelson, 1994, with Recent species from Froese & Pauly, 2001 [“The FishBase”]

<==o Salmonoidei sensu Johnson & Patterson 1997 [Salmoniformes sensu Nelson, 1994] (salmons, trouts, graylings, whitefishes and ciscoes; lohikalat)
   |?- †Goudkoffia U. Cret. NA.
   |?- †Leucichthyops U. Cret. NA.
   |?- †Natlandia U. Cret. NA.
   |?- †Salmodium Eoc. Eu.
   |-- †Beckius Eoc. NA.
   |-- †Cyclolepis U. Cret. NA.
   |-- †Cyclolepoides Eoc. NA.
   |-- †Paleolox Mioc.-Plioc. NA.
   |-- †Parastenodus Eoc. NA.
   |-- †Procharacinus Eoc. Eu.
   |-- †Prohydrocyon Eoc. Eu.
   |--o †Thaumaturus M. Eoc. L. Mioc. Eu. [Thaumaturidae]
   |  `-- †T. intermedius
   `--o Salmonidae (lohet) [FB: 12/163]
      |-- Coregoninae (whitefishes and ciscoes; siiat) [Nelson: 3 genera, 32 species] [Fishbase: 3/69]
      `--+--o Thymallinae (graylings; harjukset) [FB: 1/5]
         |  |-- †Protohymallus [Prothymallus] Mioc. Eu. 
         |  `--o Thymallus Eoc. Mioc. Eu. R. NOc. (graylings; harjukset) [Nelson: 4-5 species] [Fishbase: 5 species]
         |     |--o T. arcticus (Pallas, 1776) (Arctic grayling)
         |     |  |-- T. a. arcticus (Pallas, 1776) (Arctic grayling)
         |     |  |-- T. a. baicalensis (Dybowski, 1874) (Baikal black grayling)
         |     |  |-- T. a. grubii Dybowski, 1869 (Amur grayling)
         |     |  `-- T. a. pallasi Valenciennes, 1848 (East Siberian grayling)
         |     |-- T. brevirostris Kessler, 1879 (Mongolian grayling)
         |     |-- T. jaluensis Mori, 1928
         |     |-- T. nigrescens Dorogostaisky, 1923 (Kosogol grayling)
         |     `-- T. thymallus (Linnaeus, 1758) (harjus; Grayling)
         `-- Salmoninae (salmons and trouts; varsinaiset lohet) [FB: 8/83]

Reference(s):