After Nelson, 1994, with Recent species from Froese & Pauly, 2001 [“The Fishbase”]

<==o Mullidae (mullot; goatfishes) [N: 6/55] [FB: 6/56]
   |--o Mulloidichthys [Mulloides] [6]
   |  |-- M. dentatus (Gill, 1862) (Mexican goatfish)
   |  |-- M. flavolineatus (Lacépède, 1801) (Yellowstripe goatfish)
   |  |-- M. martinicus (Cuvier, 1829) (Yellow goatfish)
   |  |-- M. mimicus Randall & Guézé, 1980 (Mimic goatfish)
   |  |-- M. pfluegeri (Steindachner, 1900) (Orange goatfish)
   |  `-- M. vanicolensis (Valenciennes, 1831) (Yellowfin goatfish)
   |--o Mullus Mioc. Eu. R. Oc. [4]
   |  |-- †M. moldavicus
   |  |-- M. argentinae Hubbs & Marini, 1933 Argentine goatfish
   |  |-- M. auratus Jordan & Gilbert, 1882 Red goatfish
   |  |--o M. barbatus Linnaeus, 1758 (pikkumullo)
   |  |  |-- M. b. barbatus Linnaeus, 1758 (Red mullet)
   |  |  `-- M. b. ponticus Essipov, 1927
   |  `-- M. surmuletus Linnaeus, 1758 (keltajuovamullo; Striped red mullet)
   |-- Parapeneus [24]
   |--o Pseudupeneus [3]
   |  |-- P. grandisquamis (Gill, 1863) (Bigscale goatfish)
   |  |-- P. maculatus (Bloch, 1793) (Spotted goatfish)
   |  `-- P. prayensis (Cuvier, 1829) (West African goatfish)
   |--o Upeneichthys [2]
   |  |-- U. lineatus (Bloch & Schneider, 1801) (Blue-striped red mullet)
   |  `-- U. stotti Platell, Potter & Clarke, 1998
   `--o Upeneus [17]
      |-- U. arge Jordan & Evermann, 1903 (Band-tail goatfish)
      |-- U. asymmetricus Lachner, 1954 (Asymmetrical goatfish)
      |-- U. doriae (Günther, 1869) (Gilded goatfish)
      |-- U. francisi Randall & Guézé, 1992
      |-- U. japonicus (Houttuyn, 1782) (Bensasi goatfish)
      |-- U. luzonius Jordan & Seale, 1907 (Dark-barred goatfish)
      |-- U. moluccensis (Bleeker, 1855) (Goldband goatfish)
      |-- U. parvus Poey, 1852 (Dwarf goatfish)
      |-- U. pori Ben-Tuvia & Golani, 1989 (Por's goatfish)
      |-- U. quadrilineatus Cheng & Wang, 1963
      |-- U. subvittatus (Temminck & Schlegel, 1843)
      |-- U. sulphureus Cuvier, 1829 (Sulphur goatfish)
      |-- U. sundaicus (Bleeker, 1855) (Ochre-banded goatfish)
      |-- U. taeniopterus Cuvier, 1829 (Finstripe goatfish)
      |-- U. tragula Richardson, 1846 (Freckled goatfish)
      |-- U. vittatus (Forsskål, 1775) (Yellowstriped goatfish)
      `-- U. xanthogrammus Gilbert, 1892

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