After Murray, 2000, 2001, Sparks & Smith, 2004, and Kullander, 1995-2002, with species from Froese & Pauly, 2001- [“The Fishbase”]
A more complete, but non-phylogenetic list of Cichlidae -species after Froese & Pauly, 2001 ["The FishBase"]
<==o Cichlidae (kiklidit, kirjoahvenet; cichlids) |-- Etropiinae sensu Sparks & Smith, 2004 (Madagascarine and Indian cichlids; Madagaskarin ja Intian kirjoahvenet) `--+-- Ptychochrominae sensu Sparks & Smith, 2004 (ptychochromine cichlids) `--+--o Cichlinae sensu Sparks & Smith, 2004 | |--o Retroculinae sensu Kullander, 1998 (koskitonkijat) | | `--o Retroculus Eigenmann & Bray, 1894 (koskitonkijat) | | |-- R. lapidifer (Castelnau, 1855) (koskitonkija) | | |-- R. septentrionalis Gosse, 1971 | | `-- R. xinguensis Gosse, 1971 (viherkoskitonkija) | `--+--o Cichlinae sensu Kullander, 1998 (kiklat) | | |-- Cichla Bloch & Schneider, 1801 (kiklat) [Cichlini] | | `--o Crenicichlini (sukkula-ahvenet) | | |-- Crenicichla (pike cichlids; sukkula-ahvenet) | | `-- Teleocichla Kullander, 1988 (tokkoahvenet) | `--+-- Heterochromis multidens (Pellegrin, 1900) [Heterochromidinae sensu Kullander, 1998] | `--+--o Astronotinae sensu Kullander, 1998 (oskarit) | | |--o Astronotus (Oscars; oskarit, riikinkukkokirjoahvenet) [Astronotini] | | | |-- A. crassipinnis (Heckel, 1840) (paksueväoskari) | | | |-- A. ocellatus (Agassiz, 1831) (Peacock cichlid; “Oscar”; riikinkukkokirjoahven, oskari) | | | |?- A. rubrocellatus (Jardine & Schomburgk, 1843) [=? A. ocellatus] (punavatsaoskari) | | | `?- A. zebra (Pellegrin, 1904) [=? A. ocellatus] (seepraoskari) | | `--o Chaetobranchini | | |--o Chaetobranchopsis Steindachner, 1875 | | | |-- C. australis Eigenmann & Ward, 1907 | | | `-- C. orbicularis (Steindachner, 1875) | | `--o Chaetobranchus | | |-- C. flavescens Heckel, 1840 | | `-- C. semifasciatus Steindachner, 1875 | `--+--o Geophaginae sensu Kullander, 1998 (geophagines) | | |-- Geophagini | | `--+-- Crenicaratini | | `-- Acarichthyini | `--o Cichlasomatinae sensu Kullander, 1998 (acaroniine, cichlasomatine and heroine cichlids) | |-- Acaroniini (acaroniine cichlids) | `--+-- Cichlasomatini (cichlasomatine cichlids) | `-- Heroini (heroine cichlids) `--o Pseudocrenilabrinae sensu Sparks & Smith, 2004 (african cichlids; afrikankirjoahvenet) |--+-- Hemichromini | `-- Chromidotilapiini |== Tilapiini (tilapias; tilapiat) `--+?-o †Mahengechromis Murray, 2000 | |-- †M. plethos Murray, 2000 | |-- †M. rotundus Murray, 2000 | |-- †M. brachycranium Murray, 2000 | |-- †M. ellipticus Murray, 2000 | `-- †M. curvifrons Murray, 2000 |-- Cyprichromini (Lake Tanganyika herring-cichlids; Tanganjikajärven silakkakirjoahvenet) `--+--+-- Lamprologini | `-- Eretmodini |--+-- Limnochromini | `-- Ectodini `--+-- Tropheini (Lake Tanganyika mbuna-cichlids?) `-- Haplochromini (East African cichlids)
African cichlids after Dunz & Schliewen, 2013<==o Pseudocrenilabrinae |-- Heterochromini `--+-- Tylochromini `--+--o “Pelmatochromines” | |-- Pelmatochromis | `-- Pterochromis `--o “Haplotilapiines” |-- Etia nguti [Etiini] `--+-- Oreochromini `--+--o “Boreotilapiines” | |-- Gobiocichlini | `--+-- Coelotilapia joka [Coelotilapiini] | `--+-- Heterotilapia [Heterotilapiini] | `-- Coptodon [Coptodonini (sic)] `--+-- Pelmatolapia [Pelmatolapiini] `--o “Australotilapiines” |--+-- Steatocranini | `-- Tilapiini `--o “East African Radiation – EAR” |-- Boulengerochromini `--+-- Bathybatini `--+--+-- Lamprologini | `-- Eretmodini `--+-- Cyphotilapia [Cyphotilapiini] `--+-- Limnochromini `--+--+-- Benthochromis [Benthochromini] | `--+-- Perissodini | `-- Cyprichromini `--+-- Orthochromis |-- Ectodini `-- Haplochromini Reference(s):
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Reference(s):
- Dunz, A. R. & Schliewen, U. K., 2013: Molecular phylogeny and revised classification of the haplotilapiine cichlid fishes formerly referred to as “Tilapia”.
–Molecular Phylogenetics and Evolution: Vol. 68, #1, pp. 64-80 [doi: 10.1016/j.ympev.2013.03.015] - Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org - Kullander, S. O., 1995-2002: Phylogeny of major groups of cichlids. (Updated: 2002-04-12)
–iNet: Swedish Museum of Natural History: Guide to the South American Cichlidae: http://www2.nrm.se/ve/pisces/acara/cichphyl.shtml; [visited: 2005-03-11] - Murray, A. M., 2000: Eocene cichlid fishes from Tanzania, East Africa.
–Journal of Vertebrate Paleontology: Vol. 20, #4, pp. 651-664 - Murray, A. M., 2001: The fossil record and biogeography of the Cichlidae (Actinopterygii: Labroidei).
–Biological Journal of the Linnean Society: Vol. 74, #4, pp. 517-532 - Sparks, J. S., 2002: Ptychochromis inornatus; A New Cichlid (Teleostei: Cichlidae) from Northwestern Madagascar, with a Discussion of Intrageneric Variation in Ptychochromis.
–Copeia: Vol. 102, #1, pp. 120-130 - Sparks, J. S. & Smith, W. L., 2004: Phylogeny and biogeography of cichlid fishes (Teleostei: Perciformes: Cichlidae).
–Cladistics: Vol. 20, pp. 501-517