After Carroll, 1988, and Nelson, 1994, with Recent species from Froese & Pauly, 2001 [“The Fishbase”]

<==o Ophidiidae Rafinesque, 1810 [Brotulidae] (partanilkat; cusk-eels) [FB: 50/242]
   |?- †Ampheristus
   |?- †Bauzaia Eoc. NA.
   |?- †Eolamprogrammus senectus L. Eoc. WAs.
   |?-o †Propteridium L. Olig. SWAs.
   |  `-- †P. douvillei
   |?- Acanthonus armatus Günther, 1878 (Bony-eared assfish)
   |?- Alcockia rostrata (Günther, 1887)
   |?-o Apagesoma [2]
   |  |-- A. delosommatus (Hureau, Staiger & Nielsen, 1979)
   |  `-- A. edentatum Carter, 1983
   |?-o Barathrites [2]
   |  |-- B. iris Zugmayer, 1911
   |  `-- B. parri Nybelin, 1957
   |?-o Barathrodemus [2]
   |  |-- B. manatinus Goode & Bean, 1883
   |  `-- B. nasutus Smith & Radcliffe, 1913
   |?-o Bathyonus [3]
   |  |-- B. caudalis (Garman, 1899)
   |  |-- B. laticeps (Günther, 1878)
   |  `-- B. pectoralis Goode & Bean, 1885
   |?- Benthocometes robustus (Goode & Bean, 1886)
   |?- Dannevigia tusca Whitley, 1941 (Australian tusk)
   |?-o Enchelybrotula [2]
   |  |-- E. gomoni Cohen, 1982
   |  `-- E. paucidens Smith & Radcliffe, 1913
   |?- Epetriodus freddyi Cohen & Nielsen, 1978
   |?-o Holcomycteronus [6]
   |  |-- H. aequatoris (Smith & Radcliffe, 1913)
   |  |-- H. brucei (Dollo, 1906)
   |  |-- H. digittatus Garman, 1899
   |  |-- H. profundissimus (Roule, 1913)
   |  |-- H. pterotus (Alcock, 1890)
   |  `-- H. squamosus (Roule, 1916)
   |?-o Hoplobrotula [3]
   |  |-- H. armata (Temminck & Schlegel, 1846)
   |  |-- H. badia Machida, 1990
   |  `-- H. gnathopus (Regan, 1921) (False kinglip)
   |?- Hypopleuron caninum Smith & Radcliffe, 1913
   |?- Leptobrotula breviventralis Nielsen, 1986
   |?-o Leucicorus [2]
   |  |-- L. atlanticus Nielsen, 1975
   |  `-- L. lusciosus Garman, 1899
   |?-o Luciobrotula [4]
   |  |-- L. bartschi Smith & Radcliffe, 1913
   |  |-- L. corethromycter Cohen, 1964
   |  |-- L. lineata (Gosline, 1954)
   |  `-- L. nolfi Cohen, 1981
   |?-o Mastigopterus [2]
   |  |-- M. imperator Smith & Radcliffe, 1913
   |  `-- M. praetor Smith & Radcliffe, 1913
   |?- Penopus microphthalmus (Vaillant, 1888)
   |?-o Pycnocraspedum [5]
   |  |-- P. armatum Gosline, 1954
   |  |-- P. fulvum Machida, 1984
   |  |-- P. microlepis (Matsubara, 1943)
   |  |-- P. phyllosoma (Parr, 1933)
   |  `-- P. squamipinne Alcock, 1889
   |?- Selachophidium guentheri Gilchrist, 1903 (Günther's cuskeel)
   |?-o Spottobrotula [2]
   |  |-- S. amaculata Cohen & Nielsen, 1982
   |  `-- S. mahodadi Cohen & Nielsen, 1978
   |?- Tauredophidium hextii Alcock, 1890
   |?- Typhlonus nasus Günther, 1878
   |?- Xyelacyba myersi Cohen, 1961
   |--o Brotulotaenia [Brotulotaeniinae] [4]
   |  |-- B. brevicauda Cohen, 1974
   |  |-- B. crassa Parr, 1934 (Violet cuskeel)
   |  |-- B. nielseni Cohen, 1974
   |  `-- B. nigra Parr, 1933
   |-- Brotulinae [1/5]
   |-- Ophidiinae Rafinesque, 1810 (partanilkat) [8/60]
   `-- Neobythitinae [36/140]

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