After Nelson, 1994, Minckley & DeMarais., 2000, Dowling, Tibbets, Minckley & Smith, 2002, Scheurer, Bestgen & Fausch, 2003, with species from Froese & Pauly, 2001
<==o Leuciscinae [incl. Abramidinae; Chondrostominae] (särjensukuiset) |?- Aaptosyax grypus Rainboth, 1991 |?-o Algansea [6] | |-- A. aphanea Barbour & Miller, 1978 | |-- A. barbata Alvarez & Cortés, 1964 | |-- A. lacustris Steindachner, 1895 | |--o A. monticola Barbour & Contreras-Balderas, 1968 | | |-- A. m. archidion Barbour & Miller, 1994 | | |-- A. m. avia Barbour & Miller, 1978 | | `-- A. m. monticola Barbour & Contreras-Balderas, 1968 | |-- A. popoche Jordan & Snyder, 1899 | `-- A. tincella (Valenciennes, 1854) |?-o Aspius Mioc. R. Eu. [2 Recent species] | |-- †A. gracilis | |-- A. aspius (Linnaeus, 1758) (toutain; asp) | `-- A. vorax Heckel, 1843 |?- Chondrostoma Olig. R. Eu. ?Mioc. As. [26] |?-o Dionda Pleist. R. NA. [8] | |-- D. catostomops Hubbs & Miller, 1977 | |-- D. diaboli Hubbs & Brown, 1957 (Devils River minnow) | |-- D. dichroma Hubbs & Miller, 1977 | |-- D. episcopa Girard, 1856 (Roundnose minnow) | |-- D. erimyzonops Hubbs & Miller, 1977 | |-- D. ipni (Alvarez & Navarro, 1953) | |-- D. mandibularis Contreras-Balderas & Verduzco-Martinez, 1977 | `-- D. rasconis (Jordan & Snyder, 1900) |?- Elopichthys bambusa (Richardson, 1845) (Yellowcheek) |?-o Erimystax [4] | |-- E. cahni (Hubbs & Crowe, 1956) (Slender chub) | |-- E. dissimilis (Kirtland, 1840) (Streamline chub) | |-- E. insignis (Hubbs & Crowe, 1956) (Blotched chub) | `-- E. x-punctatus (Hubbs & Crowe, 1956) (Gravel chub) |?-o †Evarra | |-- †E. bustamantei Navarro, 1955 (Mexican Dace) | |-- †E. eigenmanni Woolman, 1895 | `-- †E. tlahuacensis Meek, 1902 |?-o Exoglossum | |-- E. laurae (Hubbs, 1931) (Tonguetied minnow) | `-- E. maxillingua (Lesueur, 1817) (Cutlips minnow) |?- Hesperoleucus symmetricus (Baird & Girard, 1854) (California roach) |?-o Hypophthalmichthys Plioc. R. EAs. [2] | |-- H. harmandi Sauvage, 1884 | `-- H. molitrix (Valenciennes, 1844) (Silver carp) |?- Lavinia exilicauda Baird & Girard, 1854 (Hitch) |?- Leuciscus [Idus] Olig. R. Eu. Mioc. R. As. R. Af. NA. (säyneet) [40] |?- Luciobrama macrocephalus (Lacépède, 1803) |?-o Luciocyprinus [2] | |-- L. langsoni Vaillant, 1904 | `-- L. striolatus Cui & Chu, 1986 |?-o Lythrurus [9] | |-- L. ardens (Cope, 1868) (Rosefin shiner) | |-- L. atrapiculus (Snelson, 1972) (Blacktip shiner) | |-- L. bellus (Hay, 1881) (Pretty shiner) | |-- L. fumeus (Evermann, 1892) (Ribbon shiner) | |-- L. lirus (Jordan, 1877) (Mountain shiner) | |-- L. matutinus Cope, 1870 (Pinewoods shiner) | |-- L. roseipinnis (Hay, 1885) (Cherryfin shiner) | |-- L. snelsoni (Robison, 1985) (Ouachita shiner) | `-- L. umbratilis (Girard, 1856) (Redfin shiner) |?-o Macrhybopsis | |-- M. aestivalis (Girard, 1856) (Speckled chub) | |-- M. gelida (Girard, 1856) (Sturgeon chub) | |-- M. meeki (Jordan & Evermann, 1896) (Sicklefin chub) | `-- M. storeriana (Kirtland, 1844) (Silver chub) |?-o Mylocheilus Mioc.-Plioc. R. NA. [1 Recent species] | `-- M. caurinus (Richardson, 1836) (Peamouth) |?-o Notemigonus Pleist. R. NA. [1 Recent species] | `-- N. crysoleucas (Mitchill, 1814) (Golden shiner) |?- Notropis [Hybopsis; Alburnops] Mioc. Pleist. R. NA. [N: ~70] [FB: 94] |?-o Phenacobius [5] | |-- P. catostomus Jordan, 1877 (Riffle minnow) | |-- P. crassilabrum Minckley & Craddock, 1962 (Fatlips minnow) | |-- P. mirabilis (Girard, 1856) (Suckermouth minnow) | |-- P. teretulus Cope, 1867 (Kanawha minnow) | `-- P. uranops Cope, 1867 (Stargazing minnow) |?- Platygobio gracilis (Richardson, 1836) (Flathead chub) |?-o Pogonichthys [2] | |-- †P. ciscoides Hopkirk, 1974 (Clear Lake splittail) | `-- P. macrolepidotus (Ayres, 1854) (Splittail) |?-o Pseudophoxinus [16] | |-- P. anatolicus (Hanko, 1924) | |-- P. antalyae Bogutskaya, 1992 | |-- P. battalgili Bogutskaya, 1998 | |-- P. beoticus (Stephanidis, 1939) | |-- P. callensis (Guichenot, 1850) | |-- P. crassus (Ladiges, 1960) | |-- P. drusensis (Pellegrin, 1933) | |-- P. egridiri (Karaman, 1972) | |-- P. fahirae (Ladiges, 1960) | |-- P. handlirschi (Pietschmann, 1933) | |-- P. kervillei (Pellegrin, 1911) | |-- P. maeandri (Ladiges, 1960) | |-- P. maeandricus (Ladiges, 1960) | |-- P. minutus (Karaman, 1924) | |-- P. punicus (Pellegrin, 1920) | `-- P. stymphalicus (Valenciennes, 1844) |?-o Ptychocheilus [Squalius] Mioc.-Plioc. R. NA. [4] | |-- P. grandis (Ayres, 1854) (Sacramento pikeminnow) | |-- P. lucius Girard, 1856 (Colorado pikeminnow) | |-- P. oregonensis (Richardson, 1836) (Northern pikeminnow) | `-- P. umpquae Snyder, 1908 (Umpqua pikeminnow) |?-o Richardsonius Plioc. R. NA. [2] | |-- †R. turneri | |-- R. balteatus (Richardson, 1836) (Redside shiner) | `-- R. egregius (Girard, 1858) (Lahontan redside) |?-o Rutilus Eoc. R. Eu. (roachlings; särjet) [16] | |-- R. alburnoides (Steindachner, 1866) | |-- R. arcasii (Steindachner, 1866) | |-- R. atropatenus Derjavin, 1937 (Shirvan roachling) | |-- R. aula (Bonaparte, 1841) | |-- R. basak (Heckel, 1843) | |-- R. frisii (Nordmann, 1840) (Kutum) | |-- R. karamani Fowler, 1977 | |-- R. lemmingii (Steindachner, 1866) | |-- R. lusitanicus (Collares-Pereira, 1980) | |-- R. macrolepidotus (Steindachner, 1866) | |-- R. meidingerii (Heckel, 1851) | |-- R. ohridanus (Karaman, 1924) | |-- R. pigus (Lacépède, 1803) | |-- R. rubilio (Bonaparte, 1837) | |-- R. rutilus (Linnaeus, 1758) (roach; särki) | `-- R. ylikiensis Economidis, 1991 |?-o Scardinius Olig. R. Eu. Mioc. NAs. (Rudds; sorvat) [4] | |-- S. acarnanicus Economidis, 1991 | |-- S. erythrophthalmus (Linnaeus, 1758) (sorva; Rudd) | |-- S. graecus Stephanidis, 1937 | `-- S. scardafa (Bonaparte, 1837) |?-o Telestes Bonaparte, 1837 [Leuciscus (in partim)] | |-- T. beoticus (Stephanidis, 1939) | |-- T. croaticus (Steindachner, 1866) (Croatian pijor) | |-- T. dabar Bogutskaya, Zupancic, Bogut & Naseka, 2012 | |-- T. fontinalis (M. S. Karaman (sr), 1972) (Spring pijor) | |-- T. karsticus Marcic, Buj, Duplic, Caleta, Mustafic, Zanella, Zupancic & Mrakovcic, 2011 | |-- T. metohiensis (Steindachner, 1901) | |-- T. miloradi Bogutskaya, Zupancic, Bogut & Naseka, 2012 | |-- T. montenigrinus (Vukovic, 1963) | |-- T. muticellus (Bonaparte, 1837) (Vairone) | |-- T. pleurobipunctatus (Stephanidis, 1939) | |-- T. polylepis Steindachner, 1866 | |-- T. souffia (A. Risso, 1827) (Souffia, Western Vairone) | |-- T. turskyi (Heckel, 1843) | `-- (†)T. ukliva (Heckel, 1843) |?-o Tribolodon [3] | |-- T. brandtii (Dybowski, 1872) (Pacific redfin) | |-- T. ezoe Okada & Ikeda, 1937 | `-- T. hakuensis (Günther, 1877) (Big-scaled redfin) |--o Abramidini | |--o Abramis [Ballerus] Mioc. NAs. Mioc. R. Eu. (lahnat) | | |-- A. ballerus (Linnaeus, 1758) (sulkava; Zope) | | |-- A. brama (Linnaeus, 1758) (lahna; Carp bream) | | `-- A. sapa (Pallas, 1814) (valkosilmälahna; White-eye bream) | |--o Blicca Heckel, 1843; L.Eoc.-R. Eu. Mioc.-R. NAs. (pasurit) [1 Recent species] | | `-- B. bjoerkna (Linnaeus, 1758) [Cyprinus bjoerkna Linnaeus, 1758; Abramis bjoerkna (Linnaeus, 1758)] (pasuri; White bream) | `--o Vimba (vimpat) [2] | |-- V. melanops (Heckel, 1837) (balkaninvimpa; Macedonian vimba) | `-- V. vimba (Linnaeus, 1758) (vimpa; Baltic vimba) |--+--+--o Orthodon Mioc.-Plioc. R. NA. [1 Recent species] | | | `-- O. microlepidotus (Ayres, 1854) (Sacramento blackfish) | | `--+-- Moapa coriacea Hubbs & Miller, 1948 (Moapa dace) | | `--+-- Gila Baird & Girard, 1854 [?Anchypopsis; Siphateles] Mioc. R. NA. (Chubs) | | `--o Acrocheilus Mioc.-Plioc. R. NA. [1 Recent species] | | `-- A. alutaceus Agassiz & Pickering, 1855 (Chiselmouth) | `--+--+--o Rhinichthys L. Pleist. NA. | | | |-- R. atratulus (Hermann, 1804) (Blacknose dace) | | | |-- R. cataractae (Valenciennes, 1842) (Longnose dace) | | | |-- R. chrysogaster (Girard, 1856) (Longfin dace) | | | |-- †R. deaconi Miller, 1984 (Las Vegas Dace) | | | |-- R. evermanni Snyder, 1908 (Umpqua dace) | | | |-- R. falcatus (Eigenmann & Eigenmann, 1893) (Leopard dace) | | | |-- R. osculus (Girard, 1856) (Speckled dace) | | | `-- R. umatilla (Gilbert & Evermann, 1894) (Umatilla dace) | | `--+--o Campostoma Pleist. R. NA. [] | | | |-- C. anomalum (Rafinesque, 1820) (Central stoneroller) | | | |-- C. oligolepis Hubbs & Greene, 1935 (Largescale stoneroller) | | | |-- C. ornatum Girard, 1856 (Mexican stoneroller) | | | |-- C. pauciradii Burr & Cashner, 1983 (Bluefin stoneroller) | | | `-- C. pullum (Agassiz, 1854) | | `--o Nocomis Plioc. R. NA. [7] | | |-- N. asper Lachner & Jenkins, 1971 (Redspot chub) | | |-- N. biguttatus (Kirtland, 1840) (Hornyhead chub) | | |-- N. effusus Lachner & Jenkins, 1967 (Redtail chub) | | |-- N. leptocephalus (Girard, 1856) (Bluehead chub) | | |-- N. micropogon (Cope, 1865) (River chub) | | |-- N. platyrhynchus Lachner & Jenkins, 1971 (Bigmouth chub) | | `-- N. raneyi Lachner & Jenkins, 1971 (Bull chub) | `--+-- ?Agosia [not recognized by the Fishbase] | `--+-- Cyprinella [30] | |--o Hybognathus Pleist. R. NA. [7] | | |-- H. amarus (Girard, 1856) (Rio Grande silvery minnow) | | |-- H. argyritis Girard, 1856 (Western silvery minnow) | | |-- H. hankinsoni Hubbs, in Jordan, 1929 (Brassy minnow) | | |-- H. hayi Jordan, 1885 (Cypress minnow) | | |-- H. nuchalis Agassiz, 1855 (Mississippi silvery minnow) | | |-- H. placitus Girard, 1856 (Plains minnow) | | `-- H. regius Girard, 1856 (Eastern silvery minnow) | |--o Luxilus [9] | | |-- L. albeolus (Jordan, 1889) (White shiner) | | |-- L. cardinalis (Mayden, 1988) (Cardinal shiner) | | |-- L. cerasinus (Cope, 1868) (Crescent shiner) | | |-- L. chrysocephalus Rafinesque, 1820 (Striped shiner) | | |-- L. coccogenis (Cope, 1868) (Warpaint shiner) | | |-- L. cornutus (Mitchill, 1817) (Common shiner) | | |-- L. pilsbryi (Fowler, 1904) (Duskystripe shiner) | | |-- L. zonatus (Agassiz, 1863) (Bleeding shiner) | | `-- L. zonistius Jordan, 1880 (Bandfin shiner) | `--o Pimephales Pleist. R. NA. [4] | |-- P. notatus (Rafinesque, 1820) (Bluntnose minnow) | |-- P. promelas Rafinesque, 1820 (Fathead minnow) | |-- P. tenellus (Girard, 1856) (Slim minnow) | `-- P. vigilax (Baird & Girard, 1853) (Bullhead minnow) `--+--o Semotilus L. Pleist. R. NA. [4] | |-- S. atromaculatus (Mitchill, 1818) (Creek chub) | |-- S. corporalis (Mitchill, 1817) (Fallfish) | |-- S. lumbee Snelson & Suttkus, 1978 (Sandhills chub) | `-- S. thoreauianus Jordan, 1877 (Dixie chub) `--+-- Couesius plumbeus (Agassiz, 1850) [Coesius??] (Lake chub) `--+-- Margariscus margarita (Cope, 1867) (Pearl dace) `--o Plagopterini |--+-- Meda fulgida Girard, 1856 (Spikedace) | `-- Plagopterus argentissimus Cope, 1874 (Woundfin) `--+-- Snyderichthys copei (Jordan & Gilbert, 1881) Miller, 1945 [Gila copei (Jordan & Gilbert, 1881); Gila (Snyderichthys) copei (Jordan & Gilbert) Uyeno, 1960] (Leatherside chub) [Spanish Fork & Sevier River] `--+-- Lepidomeda vittata Cope, 1874 (Little Colorado spinedace) `--+-- Snyderichthys copei [Goose Greek & Sulphur Creek] `--o Lepidomeda [4] |-- L. albivallis Miller & Hubbs, 1960 (White River spinedace) |?- †L. altivelis Miller & Hubbs, 1960 (Pahranagat spinedace) [not recognized by Dowling et al., 2002 (due extinction?)] `--o L. mollispinis Miller & Hubbs, 1960 (Virgin spinedace) |-- L. m. pratensis `-- L. m. mollispinis Miller & Hubbs, 1960
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Dowling, T. E., Tibbets, C. A., Minckley, W. L. & Smith, G. R., 2002: Evolutionary relationships of the plagopterins (Teleostei: Cyprinidae) from Cytochrome b sequences.
–Copeia: Vol. 102, #3, pp. 665-678 - Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 5: Kalat, sammakkoeläimet, matelijat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beauté Du Monde Animal: IX Reptiles/Amphibiens & X Poissons.
–Rizzoli Editore, Milano, 1968. - Minckley, W. L. & DeMarais, B. D., 2000: Taxonomy of Chubs (Teleostei, Cyprinidae, genus Gila), in the American southwest with comments on conservation.
–Copeia: Vol. 100, #1, pp. 251-256 - Nelson, J. S., 1994: Fishes of the world.
–John Wiley & Sons Inc., New York, 1994, xx-600 - Scheurer, J. A., Bestgen, K. R. & Fausch, K. D., 2003: Resolving Taxonomy and Historic Distribution for Conservation of Rare Great Plains Fishes: Hybognathus (Teleostei: Cyprinidae) in Eastern Colorado Basins.
–Copeia: Vol. 103, #1, pp. 1-12 - Zanella, D., Mrakovcic, M., Mustafic, P. & Caleta, M., 2008: Recovery of Telestes ukliva, an endemic species from the Cetina River, Croatia (Cypriniformes, Cyprinidae).
–Journal of Fish Biology: Vol. 73, #1, pp. 311-316 [doi: 10.1111/j.1095-8649.2008.01928.x]