After Nelson, 1994, with Recent species from Froese & Pauly, 2001 [“The Fishbase”]
<==o Acheilognathinae (katkerokalamaiset; bitterling-like cyprinids) |?-o Acanthorhodeus | |-- A. asmussii (Dybowski, 1872) (Russian bitterling) | |-- A. gracilis Regan, 1908 | |-- A. longibarbatus Mai, 1978 | `-- A. polyspinus Holcík, 1972 |--o Acheilognathus [Acanthorhodeus, Paracheilognathus, Pseudoperilampus] [21] | |-- A. barbatulus Günther, 1873 | |-- A. barbatus Nichols, 1926 | |-- A. chankaensis (Dybowski, 1872) (Khanka spiny bitterling) | |-- A. cyanostigma Jordan & Fowler, 1903 (Striped bitterling) | |-- A. deignani (Smith, 1945) | |-- A. elongatus (Regan, 1908) | |-- A. gracilis Nichols, 1926 | |-- A. hypselonotus (Bleeker, 1871) | |-- A. imberbis Günther, 1868 | |-- A. longipinnis (Regan, 1905) (Deepbody bitterling) | |-- A. macropterus (Bleeker, 1871) | |-- A. melanogaster Bleeker, 1860 | |-- A. meridianus (Wu, 1939) | |-- A. microphysa Yang, Chu & Chen, 1990 | |-- A. omeiensis (Shih & Tchang, 1934) | |-- A. polylepis (Wu, 1964) | |-- A. rhombeus (Temminck & Schlegel, 1846) | |-- A. tabira Jordan & Thompson, 1914 | |-- A. tonkinensis (Vaillant, 1892) | |-- A. typus (Bleeker, 1863) | `-- A. yamatsutae Mori, 1928 |?- Paracheilognathus himantegus (Günther, 1868) |--o Rhodeus Mioc. R. Eu. [14] | |-- †R. elongatus | |-- R. amarus (Bloch, 1782) | |-- R. atremius (Jordan & Thompson, 1914) (Kyushu bitterling) | |-- R. fangi (Miao, 1934) | |-- R. haradai Arai, Suzuki & Shen, 1990 | |-- R. hondae (Jordan & Metz, 1913) | |-- R. laoensis Kottelat, Doi & Musikasinthorn, 1998 | |--o R. lighti (Wu, 1931) (Light's bitterling) | | |-- R. l. lighti (Wu, 1931) (Light's bitterling) | | `-- R. l. amurensis (Vronsky, 1967) | |--o R. ocellatus (Kner, 1866) (Rosy bitterling) | | |-- R. o. ocellatus (Kner, 1866) (Rosy bitterling) | | `-- R. o. smithii (Regan, 1908) | |-- R. rheinardti (Tirant, 1883) | |-- R. sericeus (Pallas, 1776) (katkerokala; Bitterling) | |-- R. sinensis Günther, 1868 | |-- R. spinalis Oshima, 1926 | |-- R. suigensis (Mori, 1935) | `-- R. uyekii (Mori, 1935) `--o Tanakia [4] |-- T. lanceolata (Temminck & Schlegel, 1846) |-- T. limbata (Temminck & Schlegel, 1846) (Oily bitterling) |-- T. signifer (Berg, 1907) `-- T. tanago (Tanaka, 1909) (Tokyo bitterling)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Frickhinger, K. A., 1995: Fossil Atlas – Fishes.
–Mergus – Publishers for Natural History and Pet Books, Hans A. Baensch, Malle, Germany, 1-1088 - Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 5: Kalat, sammakkoeläimet, matelijat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beauté Du Monde Animal: IX Reptiles/Amphibiens & X Poissons.
–Rizzoli Editore, Milano, 1968 - Nelson, J. S., 1994: Fishes of the world.
–John Wiley & Sons Inc., New York, 1994, xx-600