Mikko's Phylogeny Archive

Nemacheilidae: genus Triplophysa Rendahl, 1933

After Kottelat, 2012

<==o Triplophysa Rendahl, 1933 [Diplophysa Kessler, 1874(preocc.), non Diplophysa Gegenbaur, 1850 (a cnidarian); Nemacheilus (Triplophysa) Rendahl, 1874; Nemacheilus (Tauphysa) Rehdahl, 1933; Deuterophysa Rendahl, 1933(preocc.), non Deuterophysa Warren, 1889 (Insecta: Lepidoptera); Didymophysa Whitley, 1950; Diplophysoides Fowler, 1958; Linemacheilus Anon. 2005 (nomen nudum)]
   |?- ?T. malloryi Banarescu & Nalbant, 1995 (nomen nudum)
   |?- ?T. akhtari (Vijayalakshmanan, 1950) [Nemacheilus akhtari Vijayalakshmanan, 1950]
   |-- T. alexandrae Prokofiev, 2001
   |-- T. aliensis Zhu & Wu in Wu & Zhu, 1979)
   |-- T. altipinnis Prokofiev, 2003
   |-- T. aluensis Li & Zhu, 2000
   |-- T. angeli (Fang, 1941) [Nemacheilus angeli Fang, 1941]
   |-- T. anterodorsalis Zhu & Cao, 1989
   |-- T. aquaecaeruleae Prokofiev, 2001
   |-- T. arnoldii Prokofiev, 2006
   |-- T. bashanensis Xu & Wang, 2009 [incl. Triplophysa longchiensis Xu & Wang, 2009; Kottelat, 2012]
   |-- T. bellibarus (Tchang, Yueh & Hwang, 1963) [Nemachilus bellibarus Tchang, Yueh & Hwang, 1963]
   |-- T. bleekeri (Sauvage & Dabry de Thiersant, 1874) [Nemachilus Bleekeri Sauvage & Dabry de Thiersant, 1874]
   |-- T. brachyptera (Herzenstein, 1888) [Nemachilus robustus brachypterus Herzenstein, 1888; ?Nemacheilus minxianensis Wang & Zhu, 1979]
   |-- T. brahui (Zugmayer, 1912) [Nemachilus brahui Zugmayer, 1912]
   |-- T. brevibarba Ding, 1993 [Triplophysa dingi Prokofiev, 2010, unjust. emend.]
   |-- T. brevicauda (Herzenstein, 1888) [Nemachilus Stoliczkae brevicauda Herzenstein, 1888]
   |-- T. cakaensis Cao & Zhu, 1988
   |-- T. chondrostoma (Herzenstein, 1888) [Nemachilus chondrostoma Herzenstein, 1888; Hedinichthys chondrostoma (Herzenstein); Wu & Wu, 1992]
   |-- T. coniptera (Turdakov, 1954) [Nemachilus conipterus Turdakov, 1954; Nemachilus conipterus salari Turdakov, 1954]
   |-- T. crassilabris Ding, 1994
   |-- T. cuneicephala (Shaw & Tchang, 1931) [Barbatula cuneicephalus Shaw & Tchang, 1931]
   |-- T. dalaica (Kessler, 1876) [Diplophysa dalaica Kessler, 1876; Nemachilus djaggasteensis rendahl, 1922; Brabatula toni posteroventralis Nichols, 1925]
   |-- T. daqiaoensis Ding, 1993
   |-- T. dorsalis (Kessler, 1872) [Cobitis dorsalis Kessler, 1872; ?N[emacheilus] (Deurterophysa) dorsalis valerii Mitrofanov, 2002 (nomen nudum)] (Grey Loach)
   |-- T. dorsonotata (Kessler, 1879) [Nemachilus dorsonotatus Kessler, 1879; Nemacheilus cuëljuensis Anikin, 1905]
   |-- T. drassensis (Tilak, 1990) [Noemacheilus drassensis Tilak, 1990)]
   |?- ?T. elegans (Kessler, 1874) [Cobitis elegans Kessler, 1874]
   |-- T. farwelli (Hora, 1935) [Nemachilus farwelli Hora, 1935]
   |-- T. flavicorpus Yang, Chen & Lan, 2004
   |-- T. furva Zhu, 1992
   |-- T. fuxianensis Yang & Chu, 1990
   |-- T. gejiuensis (Chu & Chen, 1979) [Noemacheilus gejiuensis Chu & Chen, 1979]
   |-- T. gerzeensis Cao & Zhu, 1988
   |-- T. gracilis (Day, 1877) [Nemachilus gracilis Day, 1877; ?Nemachilus deTerrai Hora, 1936; ?Triplophysa trewavasae Mirza & Ahmad, 1990]
   |-- T. grahami (Regan, 1906) [Nemachilus grahami Regan, 1906]
   |-- T. griffithi (Günther, 1868) [Nemachilus griffithii Günther, 1868; Noemacheilus naziri Ahmad & Mirza, 1963; Nemachilus griffithi hazaraensis Omer & Mirza, 1975]
   |-- T. gundriseri Prokofiev, 2002 [Nemacheilus dorsalis humilis Gundriser, 1962; Triplophysa gundriseri chandagaitensis Prokofiev, 2002]
   |-- T. hexiensis (Zhao &, Wang, 1988) [Temachilus dorsonotatus hexiensis Zhao &, Wang, 1988; ?Triplophysa rossoperegrinatorum Prokofiev, 2001]
   |-- T. heyangensis Zhu, 1992
   |-- T. hialmari Prokofiev, 2001
   |-- T. hsutschouensis (Rendahl, 1933) [Nemacheilus hsutschouensis Rendahl, 1933]
   |-- T. huanglongensis Gao, 1992
   |-- T. huanjiangensis Yang, Wu & Lan, 2011
   |-- T. huapingensis Zheng, Yang & Chen, 2012
   |-- T. hutjertjuensis (Rendahl, 1933) [Nemacheilus hutjertjuensis Rendahl, 1933)]
   |-- T. intermedia (Kessler, 1876) [Diplophysa intermedia Kessler, 1876]
   |-- T. jianchuanensis Zheng, Du, Chen & Yang, 2010 (Jianchuan Stone Loach)
   |-- T. jiarongensis Lin, Li & Song, 2012
   |-- T. kafirnigani (Turdakov, 1948) [Nemachilus dorsalis kafirnigani Turdakov, 1948]
   |-- T. kullmanni Banarescu &, Nalbant in Ladiges, 1975
   |-- T. kungessana (Kessler, 1879) [Diplophysa kungessana Kessler, 1879]
   |-- T. lacusnigri (Berg, 1928) [Nemachilus stoliczkae var. lacus-nigri Berg, 1928]
   |-- T. lacustris Yang & Chu, 1990
   |-- T. laterimaculata Li, Liu & Yang, 2007
   |-- T. laticeps Zhou & Cui, 1997
   |-- T. lihuensis Wu, Yang & Lan, 2012
   |-- T. lingyunensis (Liao & Luo in Liao, Wang & Luo, 1997) [Schistura lingyunensis Liao & Luo in Liao, Wang & Luo, 1997]
   |-- T. lixianensis He, Song & Zhang, 2008
   |-- T. longianguis Wu & Wu, 1984
   |-- T. longibarbata (Chen, Yang, Sket & Aljancic, 1998) [Paracobitis longibarbata Chen, Yang, Sket & Aljancic, 1998; Nemacheilus liboensis Ran, 2000 (nomen nudum); Nemacheilus liboensis Chen, 2000 (nomen nudum); Paracobitis longidorsalus Li, Ran & Chen in Ran, Li, & Chen, 2006 emend. Kottelat, 2012; ?Paracobitis maolanensis Li, Ran & Chen, 2006]
   |-- T. longipectoralis Zheng, Du, Chen & Yang, 2009
   |-- T. longliensis Ren, Yang & Chen, 2012
   |-- T. macrocephala Yang, Wu & Yang, 2012
   |-- T. macromaculata Yang in Chu & Chen, 1990
   |-- T. macrophthalma Zhu & Guo, 1985
   |-- T. markehenensis (Zhu & Wu, 1981) [Nemacheilus pseudoscleropterus markehenensis Zhu & Wu, 1981]
   |-- T. marmorata (Heckel, 1838) [Cobitis marmorata Heckel, 1838; Cobitis vittata Heckel, 1838; ?Nemachilus griffithii var. afghana Hora, 1935]
   |-- T. microphthalma (Liao & Wang in Liao, Wang & Luo, 1997) [Schistura dabryi microphthalmus Liao & Wang in Liao, Wang & Luo, 1997]
   |-- T. microphysus (Fang, 1935) [Nemacheilus microphysus Fang, 1935]
   |-- T. microps (Steindachner, 1866) [Cobitis microps Steindachner, 1866]
   |-- T. moquensis Ding, 1994
   |-- T. nandanensis Lan, Yang & Chen, 1995
   |-- T. nanpanjiangensis (Zhu & Cao, 1988) [Oreias dabryi nanpanjiangensis Zhu & Cao, 1988]
   |-- T. nasobarbatula Wang & Li, 2001
   |-- T. ninglangensis Wu & Wu, 1988
   |-- T. nujiangensa Chen, Cui & Yang, 2004
   |-- T. obscura Wang in Chen, Xu, Fang, Song & Wang, 1987
   |-- T. obtusirostra Wu & Wu, 1988
   |-- T. orientalis (Herzenstein, 1888) [Nemachilus kungessanus orientalis Herzenstein, 1888; Nemachilus kungessanus elongatus Herzenstein, 1888]
   |-- T. papillosolabiata (Kessler, 1879) [Diplophysa papilloso-labiata Kessler, 1879]
   |-- T. pappenheimi (Fang, 1935) [Nemacheilus pappenheimi Fang, 1935]
   |-- T. paradoxa (Turdakov, 1955) [Nemacheilus paradoxa Turdakov, 1955]
   |-- T. parva Chen, Li & Yang, 2009
   |-- T. "pedaschenkoi" (Berg, 1931) [Nemacheilus strauchi ulacholicus var. pedaschenkoi Berg, 1931] (nomen nudum)
   |-- T. polyfasciata Ding in Ding, Fang & Fang, 1996
   |-- T. pseudoscleroptera (Zhu & Wu, 1981) [Nemachilus pseudoscleroptera Zhu & Wu, 1981]
   |-- T. pseudostenura He, Zhang & Song, 2012
   |-- T. qiubeiensis Li & Yang in Li, Yang, Chen, Tao, Qi & Han, 2008
   |-- T. retropinnis (Herzenstein, 1888) [Nemachilus dorsonotatus retropinnis Herzenstein, 1888; Nemachilus dorsonotatus plagiognathus Herzenstein, 1888; ?>Nemacheilus Bertini Fang, 1941; Nemacheilus carassus Nikolski in Prokofiev, 2007 (nomen nudum); Triplophysa stoliczkae retropinnis (Herzenstein) Prokofiev, 2007]
   |-- T. robusta (Kessler, 1876) [Nemachilus robusta Kessler, 1876]
   |-- T. rosa Chen & Yang, 2005 [Linemacheilus wulong Anon., 2005 (nomen nudum); Heminoemacheilus wulongensis Wu, He, Zhen & Li, 2007 (nomen nudum)]
   |-- T. scapanognatha Prokofiev, 2007
   |-- T. scleroptera (Herzenstein, 1888) [Nemacheilus scleropterus Herzenstein, 1888]
   |-- T. sellaefer (Nichols, 1925) [Barbatula yarkandensis sellaefer Nichols, 1925]
   |-- T. sewerzowi (Nikolskii, 1938) [Nemachilus sewerzowi Nikolski, 1938] (Severtsov’s loach)
   |-- T. shaanxiensis Chen in Chen, Xu, Fang, Song & Wang, 1987
   |-- T. shehensis Menon, 1987 [Noemacheilus shehensis Tilak, 1990]
   |-- T. shilinensis Chen & Yang in Chen, Yang & Xu, 1992
   |-- T. shiyangensis (Zhao & Wang, 1983) [Nemachilus shiyangensis Zhao & Wang, 1983]
   |-- T. siluroides (Herzenstein, 1888) [Nemachilus siluroides Herzenstein, 1888]
   |-- T. stenura (Herzenstein, 1888) [Nemachilus stenurus Herzenstein, 1888; ?Nemachilus lhasae Regan, 1905]
   |-- T. stewarti (Hora, 1922) [Diplophysa stewarti Hora, 1922; Nemachilus panguri Hora, 1936; Nemachilus longianalis Ren & Wu, 1982; Nemachilus longianalis tangtianhensis Wu & Wu, 1988; Triplophysa longianalis (Ren & Wu, 1982)]
   |-- T. stolickai (Steindachner, 1866) [Cobitis Stolickai Steindachner, 1866; Triplophysa stoliczkae (Steindachner, 1866)] (Tibetan stone loach)
   |-- T. strauchii (Kessler, 1874) [Diplophysa Strauchii Kessler, 1874; Nemachilus strauchii transiens Herzenstein, 1888; ?Nemachilus strauchi zaisanicus Menshikov, 1937; ?Nemachilus ruzskyi Nakrashevich, 1948; ?N[emacheilus] (Deuterophysa) strauchi jimbeyi Mitrofanov, 2002] (Spotted thicklip loach)
   |-- T. tanggulaensis (Zhu, 1982) [nemacheilus tanggulaensis Zhu, 1982]
   |-- T. tianeensis Chen, Cui & Yang, 2004
   |-- T. tibetana (Regan, 1905) [Nemachilus tibetanus Regan, 1905]
   |-- T. turpanensis Wu & Wu, 1992
   |-- T. ulacholica (Anikin, 1905) [Nemachilus ulacholicus Anikin, 1905; Nemachilus strauchi dorsaloides Turdakov, 1947; Nemacheilus strauchi reuniens Turdakov, 1952; Nemacheilus strauchi ulacholicus Anikin, 1905] (Issyk-kul naked loach)
   |-- T. uranoscopus (Kessler, 1872) [Cobitis uranoscopus Kessler, 1872; Nemachilus fedtschenkoae Nikolski, 1903]
   |-- T. venusta Zhu & Cao, 1988
   |-- T. waisihani Cao & Zhang, 2008
   |-- T. wuweiensis (Li & Chang, 1974) [Nemachilus wuweiensis Li & Chang, 1974]
   |-- T. xiangshuingensis Li, 2004
   |-- T. xiangxiensis (Yang, Yuan & Liao, 1986) [Noemacheilus xiangxiensis Yang, Yuan & Liao, 1986]
   |-- T. xichangensis Zhu & Cao, 1989
   |-- T. xingshanensis (Yang & Xie, 1983) [Nemachilus xingshanensis Yang & Xie, 1983]
   |-- T. xiqiensis Ding & Lai, 1996
   |-- T. yaopeizhii Xu, Zhang & Cai in Zhang, Cai & Xu, 1995
   |-- T. yunnanensis Yang, 1990
   |-- T. zhaoi Prokofiev, 2006 [Nemachilus turfanensis Herzenstein, 1899 (nomen nudum)]
   `-- T. zhenfengensis Wang & Li, 2001