After lechtová, Bohlen & Perdices, 2008 (fig. 4 & 5), with Recent species from Froese & Pauly, 2001 [“The Fishbase”] and Kottelat, 2012
<==o Cobitidae Swainson, 1838 [Cobitidae Swainson, 1838; Acanthopsides Heckel & Kner, 1858; Misgurninae Fowler, 1905] (cobitine loaches; piikkinuoliaiset; “rantaneulat”) |--o Lepidocephalus Bleeker, 1858 | |?- L. caudofurcatus Tilak & Husain, 1978 (Tilak loach) | |-- L. macrochir (Bleeker, 1854) [Cobitis macrochir Bleeker, 1854; Lepidocephalichthys pallens Vaillant, 1902; Acanthophthalmus pahangensis de Beaufort, 1933] | `?- L. spectrum Roberts, 1989 |--+-- Neoeucirrhichthys maydelli Banarescu & Nalbant, 1968 (Goalpara loach) | `-- Canthophrys gongota (Hamilton, 1822) Swainson, 1838 [Cobitis gongota Hamilton, 1822; Cobitis cucura Hamilton, 1822; Cobitis oculata M'Clelland, 1839; Canthophrys albescens Swainson, 1838; Somileptus bispinosa Swainson, 1838; ?Cobitis amnicola Valenciennes, in Cuvier & Valenciennes, 1846] (Gongota loach) |--o Lepidocephalichthys Bleeker, 1863 [taxonomic history] | |?- L. alkaia Havird & Page, 2010 | |?- L. annandalei Chaudhuri, 1912 [taxonomic history] (Annandale loach) | |?- ?L. arunachalensis (Datta & Barman, 1984) [taxonomic history] | |?- L. barbatuloides (Bleeker, 1851) [taxonomic history] | |?- ?L. birmanicus (Rendahl, 1948) | |?- L. coromandelensis (Menon, 1992) [taxonomic history] | |?- L. furcatus (de Beaufort, 1933) [taxonomic history] | |?- ?L. goalparensis Pillai & Yazdani, 1976 [taxonomic history] (Pillai loach) | |?- L. irrorata Hora, 1921 [taxonomic history] (Loktak loach) | |?- L. jonklaasi (Deraniyagala, 1956) [taxonomic history] (Jonklaas's loach) | |?- L. kranos Havird & Page, 2010 | |?- L. lorentzi (Weber & de Beaufort, 1916) [taxonomic history] | |?- L. micropogon (Blyth, 1860) [taxonomic history] | |?- L. tomaculum Kottelat & Lim, 1992 | |?- L. zeppelini Havird & Tangjitjaroen, in Havind, Page, Tangjitjaroen, Vidthayanon, Grudpan & Udduang, 2010 | |-- L. thermalis (Valenciennes, 1846) [taxonomic history] (Common spiny loach; täplänuoliainen) | `--+-- L. hasselti (Valenciennes, 1846) [taxonomic history] | `--+-- L. guntea (Hamilton, 1822) (Guntea loach) | `-- L. berdmorei (Blyth, 1860) [taxonomic history] (Burmese loach) `?-+--+--o Acantopsis van Hasselt, 1823 [Acanthopsis van Hasselt, 1824; ?Aperioptus Richardson, 1848; Prostheacanthus Blyth, 1860] (kärsänuoliaiset) | | |-- A. dialuzona Van Hasselt, 1823 [Acantopsis Dialuzona Van Hasselt, 1823; ?Aperioptus pictorius Richardson, 1848; Acantopsis biaculeata Rüppell, 1852; Cobitis choirorhynchos Bleeker, 1854; Cobitis macrorhynchos Bleeker, 1854] (Horseface loach; kärsänuoliainen, hevosnuoliainen) | | |-- A. octoactinotos Siebert, 1991 | | |-- A. spectabilis Blyth, 1860 [Prostheacanthus spectabilis Blyth, 1860; ?Acantopsis multistigmatus Vishwanath & Laisram, 2005] | | `-- A. thiemmedhi Sontirat, 1999 [A. thiemmedhi Sontirat, 1997 (nomen nudum)] | `==o Acanthopsoides Fowler, 1934 [Neacanthopsis Smith, 1945] | |?- A. delphax Siebert, 1991 | |?- A. gracilis Fowler, 1934 (pikkukärsänuoliainen) | |?- A. molobrion Siebert, 1991 | |--+-- A. gracilentus (Smith, 1945) [Neacanthopsis gracilentus Smith, 1945; Acanthopsoides namromensis Nguyen, 2005] | | `-- A. hapalias Siebert, 1991 | `--+-- A. robertsi Siebert, 1991 | `--o Kottelatlimia Nalbant, 1994 [Lepidocephalichthys Bleeker, 1863 in partim] | |?- K. hipporhynchos Kottelat & Tan, 2008 | |-- K. katik (Kottelat & Lim, 1993) [Lepidocephalichthys katik Kottelat & Lim, 1993] | `-- K. pristes(Roberts, 1989) [Lepidocephalichthys pristes Roberts, 1989] `?-+-- Pangio Blyth, 1860 [incl. Eucirrichthys Perugia, 1892; Cobitophis Myers, 1927] [taxonomic history] (coolie-loaches; piikkisilmät) `--+-- Microcobitis misgurnoides (Rendahl, 1944) Bohlen & Harant, 2011 [Cobitis misgurnoides Rendahl, 1944; ?Cobitis nuicocensis Nguyen & Vo, in Nguyen, 2005] `?-+--o Sabanejewia Vladykov, 1829 | |?- S. balcanica (Karaman, 1922) [Cobitis balcanica Karaman, 1922; Cobitis montana Vladykov, 1925; Cobitis aurata balcanica natio radnensis Jaszfalusi, 1951; Cobitis aurata balcanica morpha elongata Oliva, Balon & Franck, 1952; Cobitis aurata balcanica morpha elata Oliva, Balon & Franck, 1952; Cobitis aurata radnensis Banarescu, Müller & Nalbant, 1960; Cobitis aurata bosniaca Karaman, 1963; ?Sabanejewia aurata doiranica Economidis, 1995 (nomen nudum); ?Sabanejewia aurata thrakica Economidis, 1995 (nomen nudum); ?Sabanejewia aurata doiranica Economidis & Nalbant, 1997; Sabanejewia aurata thrakica Economidis & Nalbant, 1997] | |?- S. baltica Witkowski, 1994 [S. aurata baltica Witkowski, 1994] | |?- S. caspia (Eichwald, 1838) [Cobitis caspia Eichwald, 1838] (Caspian spined loach) | |?- S. caucasica (Berg, 1906) [Cobitis caucasica Berg, 1906] | |?- S. kubanica Vasil'eva & Vasil'ev, 1988 [S. aurata kubanica Vasil'eva & Vasil'ev, 1988] | |?- S. vallachica (Nalbant, 1957) [Cobitis aurata vallachica Nalbant, 1957] | |-- S. larvata (De Filippi, 1859) [Cobitis larvata De Filippi, 1859; Cobitis taenia var. conspersa Cantoni, 1882] | `--+-- S. romanica (Bacescu, 1943) [Cobitis caspia romanica Bacescu, 1943] | `--+-- S. bulgarica (Drensky, 1928) [Cobitis bulgarica Drensky, 1928; Cobitis albicoloris Chichkoff, 1932; Cobitis taenia tesselatus Pietschmann, 1937] | `--o S. aurata (De Filippi, 1863) [Cobitis aurata De Filippi, 1863; Cobitis hohenackeri Kessler, 1877; ?Cobitis aralensis Kessler, 1877] (Golden spined loach) | |-- S. a. aurata (De Filippi, 1863) | `-- S. a. aralensis (Kessler, 1877) [?Cobitis aralensis Kessler, 1877] (Aral spined loach) `--+--+--o Koreocobitis Kim, Park & Nalbant, 1997 [Cobitis in partim] | | |-- K. rotundicaudata (Wakiya & Mori, 1929) [Cobitis rotundicaudata Wakiya & Mori, 1929] | | `-- K. naktongensis Kim, Park & Nalbant, 2000 | `--+-- Paramisgurnus dabryanus Dabry de Thiersant, 1872 [Paramisgurnus dabryanus Sauvage, 1878; Misgurnus mizolepis Günther, 1888;Misgurnus oligolepos Chen, Shen & Li, 1994] | `--+-- Misgurnus fossilis (Linnaeus, 1758) [taxonomic history] (euroopanmutakala, barometrikala; Weatherfish) | `--+-- Misgurnus sp.1 | `-- Misgurnus anguillicaudatus (Cantor, 1842) A [taxonomic history] (aasianmutakala, “kiinanmonni”; Oriental weatherfish) [polyphyletic] `--+--+--+-- Iksookimia choii (Kim & Son, 1984) (Choi's spiny loach) | | `--+-- Cobitis melanoleuca Nichols, 1925 [Cobitis taenia melanoleuca Nichols, 1925] | | |?- Cobitis gladkovi Vasil'eva & Vasil'ev, 2008 [Cobitis melanoleuca gladkovi Vasil'eva & Vasil'ev, 2008] | | `?- Cobitis sibirica Gladkov, 1935 [Cobitis taenia sibirica Gladkov, 1935; Cobitis taenia granoei Rendahl, 1935; Cobitis taenia sibirica morpha elongata Berg, 1949; Cobitis granoei olivai Nalbant, Holcik & Pivnicka, 1970; Cobitis melanoleuca granoei Vasil'eva & Vasil'ev, 2008] (Siberian spiny loach) | `--+-- Niwaella delicata (Niwa, 1937) Nalbant, 1963 [Cobitis delicata Niwa, 1937] [Niwaëlla Nalbant, 1963] [polyphyletic] | `--+-- Cobitis striata Ikeda, 1936 [Cobitis taenia striata Ikeda, 1936] | `-- Cobitis biwae Jordan & Snyder, 1901 [Cobitis taenia japonica Temminck & Schlegel, 1846; Cobitis Schlegeli Bleeker, 1863 (nomen oblitum)] `--+--+-- Iksookimia pacifica (Kim, Park & Nalbant, 1999) [Cobitis pacifica Kim, Park & Nalbant, 1999] | `--+-- Iksookimia pumila (Kim & Lee, 1987) [Cobitis koreensis pumilus Kim & Lee, 1987] | `-- Iksookimia koreensis (Kim, 1975) [Cobitis koreensis Kim, 1975] `--+--+-- ?Niwaella sp. cf. N. laterimaculata | |?- ?Niwaella laterimaculata (Yan & Zheng, 1984) [Cobitis laterimaculata Yan & Zheng, 1984] | |?- ?Niwaella longibarba Chen & Chen, 2005 | `?- ?Niwaella xinjiangensis Chen & Chen, 2005 `?-+--+-- Cobitis taenia Linnaeus, 1758 [Cobitis Taenia Linnaeus, 1758; Cobitis spilura Holandre, 1837; Cobitis taenioides Bacescu & Maier, 1969; Cobitis taenia taenia] (rantaneula; Spined loach) | `?-+-- Cobitis elazigensis Coad & Sarieyyüpoglu, 1988 [=? C. simplicispina Hankó, 1924 (Erkakan et al., 1999)] | `--o Cobitis (Iberocobitis) Bacescu, 1962 | |-- C. (I.) maroccana Pellegrin, 1929 [Cobitis taenia var. maroccana Pellegrin, 1929] | |-- C. (I.) paludica (de Buen, 1930) [Acanthopsis taenia forma paludica de Buen, 1930; Cobitis taenia paludicola Berg, 1932; Cobitis taenia haasi Klausewitz, 1955; ?Cobitis vitoriae Fernandez de la Cigoña Núñez & Ferreira, 1996; ?Cobitis victoriae Fernandez de la Cigoña Núñez, 1999] | `-- C. (I.) vettonica Doadrio & Perdices, 1997 `--+-- Cobitis rara Chen , 1981 `?-+--+-- Misgurnus anguillicaudatus (Cantor, 1842) B [taxonomic history] (Oriental weatherfish; aasianmutakala, “kiinanmonni”) [polyphyletic] | `--+-- Misgurnus sp.2 | `--+-- Misgurnus sp.3 | `-- Misgurnus mohoity [incl. Misgurnus bipartitus sensu Kottelat, 2012] [taxonomic history] `?-+?-+?- Cobitis bilineata Canestrini, 1866 [Cobitis taenia var. bilineata Canestrini, 1866; Cobitis taenia var. puta Cantoni, 1882; Cobitis taenia var. septa Cantoni, 1882] | `?- Cobitis elongata Heckel & Kner, 1858 `--+-- Cobitis lutheri Rendahl, 1935 [Cobitis taenia lutheri Rendahl, 1935] (Luther's spiny loach) `--+--+-- Iksookimia yongdokensis Kim & Park, 1997 | `-- Iksookimia longicorpus (Kim, Choi & Nalbant, 1976) [Cobitis longicorpus (Kim, Choi & Nalbant, 1976] `?-o Kichulchoia Kim, Park & Nalbant, 1999 [Choia Kim, Park & Nalbant, 1997 non Choia Walcott, 1920 (a fossil sponge (Porifera))] |-- K. brevifasciata Kim & Lee, 1995 [Niwaella brevifasciata Kim & Lee, 1995] `-- K. multifasciata (Wakiya & Mori, 1929) [Cobitis multifasciata Wakiya & Mori, 1929; Niwaella multifasciata (Wakiya & Mori, 1929)]
Untreated Cobitids
Bibarba bibarba Chen & Chen, 2007
Iksookimia Nalbant, 1993
I. hugowolfeldi Nalbant, 1993
Misgurnus Lacepčdé [La Cepčde], 1803 [taxonomic history] (mutakalat) [polyphyletic]
M. buphoensis Kim & Pak, 1995 [taxonomic history]
M. multimaculatus Rendahl, 1944 [taxonomic history]
M. nikolskyi Vasil'eva, 2001 [taxonomic history]
M. tonkinensis Rendahl, 1937 [taxonomic history]
Theriodes sandakanensis (Inger & Kong, 1962) Kottelat, 2012 [Acanthophthalmus sandakanensis Inger & Kong, 1962; Lepidocephalichthys sandakanensis (Inger & Kong, 1962); Pangio? sandakanensis (Inger & Kong, 1962) sensu Havird & Page, 2010]
Paralepidocephalus Tchang, 1935
?P. guishanensis Li, 2004 [Paralepidcephalus [sic] guishanensis Li, 2004]
P. yui Tchang, 1935
'Protocobitis' polylepis Zhu, Lü, Yamng & Zhang, 2008; Kottelat, 2012
Protocobitis typhlops Yang & Chen, 1993 [Protocobitis typhlops Yang, Chen & Lan, 1994]
Cobitis Linnaeus, 1758 [Acantophthalmus van Hasselt, 1823; Acanthophthalmus van Hasselt, 1823; ?Acanthopsis Agassiz, 1832; Cobitinula Hankó, 1924; Acanestrinia Bacescu, 1962; Bicanestrina Bacescu, 1962; Iberocobitis Bacescu, 1962; Beyshehiria Erk'akan, Atalay-Ekmekçi & Nalbant, 1999] Olig. R. Eu. R. As. NAf.
†G. centrochir
C. amphilekta Vasil'eva & Vasil'ev, 2012
C. arachthosensis Economidis & Nalbant, 1996 [Cobitis hellenica arahthosensis Economidis, 1991 (nomen nudum); Cobitis arachthosensis Economidis & Nalbant, 1996]
C. bilseli Battalgil, 1942
C. calderoni Bacescu, 1962 [Cobitis Calderóni Bacescu, 1962]
C. choii Kim & Son, 1984
?C. caucasica Berg, 1906 (Ciscaucasian spined loach)
?C. conspersa Cantoni, 1882
C. dalmatina Karaman, 1928 [Cobitis taenia dalmatina Karaman, 1928]
C. dolichorhynchus Nichols, 1918
C. elongatoides Bacescu & Maier, 1969 [Cobitis taenia var. elongatoďdes Bacescu, 1962; Cobitis taenia elongatoides Bacescu & Maier, 1969; Cobitis taenia danubialis Nalbant, 1993; Cobitis megaspila Nalbant, 1993]
C. evreni Erk'akan, Özeren & Nalbant, 2008
C. fahifeae Erk'akan, Atalay-Ekmekçi & Nalbant, 2008 [Cobitis fahifae Erk'akan, Atalay-Ekmekçi & Nalbant, 2008 (lapsus calami)]
C. faridpaki Mousevi-Sabet, Vasil'eva, Vatandoust & Vasil'ev, 2011
?C. haasi Klausewitz, 1955
C. hankugensis Kim, Park, Son & Nalbant, 2003
C. hellenica Economidis & Nalbant, 1997 [Cobitis hellenica Economidis, 1991 (nomen nudum)]
C. illyrica Freyhof & Stelbrink, 2007
C. jadovaensis Mustafic & Mrakovcic, in Mustafic, Marcic, Dublic, Mrakovcic, Caleta, Zanella, Buj, Podnar & Dolenec, 2008
C. kellei Erk'akan, Atalay-Ekmekçi & Nalbant, 1998
C. keyvani Mousavi-Sabet, Yerli, Vatandoust, Özeren & Moradkhani, 2012
C. laoensis (Sauvage, 1878) [Misgurnus laoensis Sauvage, 1878; ?Cobitis longitaeniatus Ngo, 2008; ?Cobitis longitaenia Ngo, 2008 (lapsus calami); ?Cobitis phongnhaensis Ngo, 2008]
?C. lebedevi Vasil'eva & Vasil'ev, 1985 [=? Cobitis choii Kim & Son, 1984 (Bogutskaya et al., 2008)]
C. levantina Krupp & Moubayed, 1992
C. linea (Heckel, 1846-49) [Acanthopsis linea Heckel, 1846-49]
C. macrostigma Dabry de Thiersant, 1872
C. matsubarae Okada & Ikeda, 1939 [Cobitis taenia matsubarae Okada & Ikeda, 1939]
C. meridionalis Karaman, 1924 [Cobitis taenia meridionalis Karaman, 1924]
C. microcephala Chen & Chen, 2011
C. multimaculata Chen & Chen, 2011
C. narentana Karaman, 1928 [Cobitis taenia narentana Karaman, 1928]
C. ohridana Karaman, 1928 [Cobitis taenia ohridana Karaman, 1928]
C. phrygica Battalgazi, 1944 [Cobitis Battalgili Bacescu, 1962] [=? Cobitis simplicispina Hankó, 1924 (Erkakan et al., 1999)]
C. pontica Vasil'eva & Vasil'ev, 2006
C. puncticulata Erk'akan, Atalay-Ekmekçi & Nalbant, 1998
C. punctilineata Economidis & Nalbant, 1997 [C. punctilineata Economidis, 1991 (nomen nudum)]
C. satunini Gladkov, 1985 [Cobitis taenia satunini Gladkov, 1985]
C. shikokuensis suzawa, 2006
C. simplicispina Hankó, 1924
C. sinensis Sauvage & Dabry de Thiersant, 1874
C. splendens Erk'akan, Atalay-Ekmekçi & Nalbant, 1998
C. stephanidisi Economidis, 1992 [C. stephanidisi Economidis, 1991 (nomen nudum); C. stephanidisi Economidis & Nalbant, 1997]
C. strumicae Karaman, 1955 [Cobitis taenia strumicae Karaman, 1955; Cobitis peschevi Sivkov & Dobrovolov, 1984; Cobitis rhodopensis Vassilev, 1998]
C. takatsuensis Mizuno, 1970
C. tanaitica Bacescu & Maier, 1969 [Cobitis taenia tanaitica Bacescu & Maier, 1969; Cobitis rossomeridionalis Vasil'eva & Vasil'ev, 1998]
C. taurica Vasil'eva, Vasil'ev, Janko, Ráb & Rábová, in Janko, Vasil'ev, Ráb, Rábová, lechtová & Vasil'eva, 2005
C. tetralineata Kim, Park & Nalbant, 1999
C. trichonica Stephanidis, 1974
C. turcica Hankó, 1924 [Cobitis taenia turcica Hankó, 1924; ?Cobitinula anatoliae Hankó, 1924]
C. vardarensis Karaman, 1928 [Cobitis taenia vardarensis Karaman, 1928; C. vardarensis kurui Erk'akan, Atalay-Ekmekçi & Nalbant, 1998]
C. ylengensis Ngo, 2003 [?Cobitis squataeniatus Ngo, 2008]
C. zanandreai Cavicchioli, 1965 [Cobitis taenia zanandreai Cavicchioli, 1965]
C. zhejiangensis Son & He, 2005
?C. arenae (Lin, 1934) [Misgurnus aranae Lin, 1934] (incertae sedis)
?C. guttata (Nguyen, 2005) [Acantopsis guttatus Nguyen, 2005] [non Cobitis guttata M'Clelland, 1839] (incertae sedis)
?C. longimanus Dybowski, in Sinicyn, 1900 (nomen nudum)
?C. lachnostoma (Rutter, 1897) [Acanthopsis lachnostoma Rutter, 1897]
?C. poecilopleura Bleeker, 1872 (nomen nudum)
?C. yeni Nguyen, 1982 (nomen nudum)
Alternative phylogenies of Cobitidae
After Ramaswami, 1953 [via lechtová, Bohlen & Perdices, 2008]
<==o Cobitidae
|--+-- Cobitis
| |-- Misgurnus
| `-- Canthophrys
`--+-- Acantopsis
|-- Pangio
`-- Lepidocephalus
|
After Nalbant, 1963 [via lechtová, Bohlen & Perdices, 2008]
<==o Cobitidae
|-- Enobarbus
`--+-- Misgurnus
|--+-- Canthophrys
| `--+-- Niwaella
| `--+-- Sabanejewia
| `-- Cobitis
`--+--+-- Acantopsis
| `-- Acantopsoides
`--+--+-- Paralepidocephalus
| `--+-- Lepidocephalichthys
| `-- Lepidocephalus
`--+-- Eucirrichthys
`--+-- Cobitophis
`-- Pangio
|
After Sawada, 1982 [via lechtová, Bohlen & Perdices, 2008]
<==o Cobitidae
|--+-- Misgurnus
| `-- Pangio
`--+-- Acantopsis
`--+-- Niwaella
`--+-- Acantopsoides
`--+-- Cobitis
|-- Iksookimia
|-- Lepidocephalichthys
`-- Sabanejewia
|
After Nalbant, 1994 [via lechtová, Bohlen & Perdices, 2008]
<==o Cobitidae
|-- Enobarbus
|-- Canthophrys
|-- Kottelatiimia
`--+--+-- Misgurnus
| |--+-- Acantopsis
| | `-- Acantopsoides
| `--+-- Cobitis
| |-- Iksookimia
| |-- Niwaella
| |-- Paralepidocephalus
| `-- Sabanejewia
`--+--+-- Eucirrichthys
| `-- Pangio
`--+-- Neoeucirrichthys
`--+-- Lepidocephalichthys
`-- Lepidocephalus
|
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Frickhinger, K. A., 1995: Fossil Atlas – Fishes.
–Mergus – Publishers for Natural History and Pet Books, Hans A. Baensch, Malle, Germany, 1-1088 - Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org - Havird, J. C. & Page, L. M., 2010: A revision of Lepidocephalichthys (Teleostei: Cobitidae) with descriptions of two new species from Thailand, Laos, Vietnam, and Myanmar.
–Copeia: Vol. 2010, #1, pp. 137-159 [doi: 10.1643/CI-08-240] - Kottelat, M., 2012: Conspectus Cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei).
–The Raffles Bulletin of Zoology: Supplement Number 26, pp. 1-199 - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 5: Kalat, sammakkoeläimet, matelijat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original publication [in French]: Beauté Du Monde Animal: IX Reptiles/Amphibiens & X Poissons.
–Rizzoli Editore, Milano, 1968 - Long, J. A., 1995: The Rise of Fishes: 500 million years of evolution.
–Johns Hopkins University Press, Baltimore & London, pp. 1-223 - Nelson, J. S., 1994: Fishes of the world.
–John Wiley & Sons Inc., New York, 1994, xx-600 - lechtová, V., Bohlen, J. & Perdices, A., 2008: Molecular phylogeny of the freshwater fish family Cobitidae (Cypriniformes: Teleostei): Delimitation of genera, mitochondrial introgression and evolution of sexual dimorphism.
–Molecular Phylogenetics and Evolution: Vol. 47, #2, pp. 812-831 [doi: 10.1016/j.ympev.2007.12.018] - Varjo, M., 2001: Akvaario-opas. [in Finnish]
–AquaGeo & Gummerus Kirjapaino Oy, Jyväskylä, 2001, 1-143