After Carroll, 1988, and Nelson, 1994, with Recent species from Froese & Pauly, 2001 [“The Fishbase”]

<==o Holocentrinae (piikkioravakalat) [3/32]
   |--o Holocentrus Eoc. Mioc. Eu. R. Oc. [N: 2] [FB: 4]
   |  |-- †H. sp. [Eocene, Monte Bolca, Italy]
   |  |-- H. ascensionis (Osbeck, 1765) (Squirrelfish)
   |  |-- H. bullisi Woods, 1955 (Deepwater squirrelfish)
   |  |-- H. coruscus Poey, 1860 (Reef squirrelfish)
   |  `-- H. rufus (Walbaum, 1792) (Longspine squirrelfish)
   |--o Neoniphon [Flammeo] [5]
   |  |-- N. argenteus (Valenciennes, 1831) Clearfin squirrelfish
   |  |-- N. aurolineatus (Liénard, 1839) Yellowstriped squirrelfish
   |  |-- N. marianus (Cuvier, 1829) Longjaw squirrelfish
   |  |-- N. opercularis (Valenciennes, 1831) [Holocentrus opercularis] (oravakala; Blackfin squirrelfish)
   |  `-- N. sammara (Forsskål, 1775) Sammara squirrelfish
   `-- Sarcocentron [Adioryx] [N: 25] [FB: 32]

Reference(s):