`--o SIPUNCULA Rafinesque, 1814 [SIPUNCULIDA Rafinesque, 1814; Sifunculacei Delle Chiaje 1823; Gephyrea de | Quatrefage 1847; Sipunculacea Hatschek 1881; Podaxonia Lankester 1885; | Prosopygia Lang 1888; Sipunculoidea Sedgwick 1898] (ruiskumadot) |--o SIPUNCULIDEA Rafinesque, 1814 | |--o SIPUNCULIFORMES Gibbs & Cutler, 1985 | | `--o Sipunculidae Rafinesque 1814 | | |--o Sipunculus Linnaeus, 1766 [Syrinx Bohadesh, 1761; Lumbricus Linnaeus, 1758] | | | |--o {Sipunculus} Linnaeus, 1766 | | | | |--- S. {S.} lomonossovi Murina, 1968 | | | | |--- S. {S.} longipapillosus Murina, 1968 | | | | |--- S. {S.} marcusi Ditadi, 1976 [?Lumbricus phalloides Pallas, 1774] | | | | |--- S. {S.} norvegicus Danielssen, 1868 [Sipunculus aequabilis Sluiter, 1902; Sipunculus | | | | | priapuloides americanua Gerould, 1913; Sipunculus infrons Sluiter, 1902; Sipunculus | | | | | nitidus Sluiter, 1900; Sipunculus priapuloides Koren & Danielssen, 1877] | | | | |--- S. {S.} nudus Linnaeus, 1766 [Syrinx tesselatus Rafinesque, 1814; Sipunculus balanophorus Delle | | | | | Chiaje, 1823; Sipunculus gigas de Quatrefages, 1856; Sipunculus eximioclathrathus | | | | | Baird, 1868; Sipunculus titubans Selenka & Balow, 1883; Sipunculus titubans | | | | | diptychus Fischer, 1895; Sipunculus delphinus Murina, 1967] | | | | |--o S. {S.} phalloides (Pallas, 1774) [Lumbricus phalloides Pallas, 1774] | | | | | |--- S. {S.} p. phalloides (Pallas, 1774) [Lumbricus phalloides Pallas, 1774; Sipunculus | | | | | | galapagensis Fischer, 1947; Sipunculus multisulcatus Fischer, 1913] | | | | | `--- S. {S.} p. inclusus Sluiter, 1902 [Sipunculus sp. Cutler & Cutler, 1985] | | | | |--- S. {S.} polymyotus Fischer, 1947 [Sipunculus natans Fischer, 1954] | | | | `--- S. {S.} robustus Keferstein, 1865 [Sipunculus angasii Baird, 1868; Sipunculus gravieri Hrubel, | | | | 1904] | | | `--o {Austrosiphon} Fischer, 1954 [Xenosiphon {Austrosiphon} Fischer, 1954; Xenosiphon {Xenopsis} | | | | Johnson, 1969; Sipunculus {Contraporus} Cutler & Cutler, 1985] | | | |--- S. {A.} indicus Peters, 1850 [Sipunculus porrectus Selenka, 1888; Sipunculus discrepans Sluiter, | | | | 1898] | | | `--- S. {A.} mundanus (Selenka & Balow, 1883) [Xenosiphon mundatus Selenka & Balow, 1883; Sipunculus | | | maoricus Benham, 1904] | | |--o Xenosiphon Fischer, 1947 | | | |--- X. absconditus Saiz, 1984 [Xenosiphon branchiatus nudud Cutler & Cutler, 1985] | | | `--- X. branchiatus (Fischer, 1895) [Sipunculus mundanus branchiatus Fischer, 1895; Xenosiphon | | | caribaceus Fischer, 1954] | | |--o Siphonosoma Spengel, 1912 [Siphonosoma {Damosiphon} Fischer, 1950; Siphonosoma {Hesperosiphon} | | | | Fischer, 1950] | | | |--- S. arcassonense Cuenot, 1902 | | | |--o S. australe (Keferstein, 1865) [Phascolosoma australe Keferstein, 1865] | | | | |--- S. s. australe (Keferstein, 1865) [Phascolosoma australe Keferstein, 1865; Siphonosoma aeneus | | | | | Baird, 1868] | | | | `--- S. takatsukii Sati, 1939 [Siphonosoma pescadolense Sati, 1939] | | | |--- S. boholense (Selenka, de Man & Balow, 1883) [Sipunculus boholense Selenka, de Man & Balow, 1883] | | | |--- S. cumanense (Keferstein, 1867) [Phascolosoma cumanense Keferstein, 1867; Lumbricus edulis Pallas, | | | | 1774; Sipunculus deformis Baird, 1868; Sipunculus cumanensis vitreus Selenka & Balow, | | | | 1883; Phascolosoma semirugosum Grabe, 1868; Sipunculus billitonensis Sluiter, 1886; | | | | Sipunculus claviger Sluiter, 1902; Sipunculus novaepommeraniae Fischer, 1926; | | | | Physcosoma hebes Sluiter, 1902; Siphonosoma carolinense Fischer, 1928; Siphonosoma | | | | hataii Sati, 1935; Siphonosoma cumanense yapense Sati, 1935; Siphonosoma formosum | | | | Sati, 1939; Siphonosoma koreae Sati, 1939; Siphonosoma opacum (Selenka & Balow, 1883); | | | | Siphonosoma marchadi Stephen, 1960] | | | |--- S. dayi Stpehen, 1942 | | | |--- S. funafuti (Shipley, 1898) [Sipunculus funafuti Shipley, 1898; Sipunculus amamiense Ikeda, 1904] | | | |--- S. ingens (Fischer, 1947) [Sipunculus ingens Fischer, 1947] | | | |--- S. mourense Sati, 1930 | | | |--- S. rotumanum (Shipley, 1898) [Sipunculus rotumanum Shipley, 1898; Siphonosoma eniwetoki Fischer, | | | | 1950; Siphonosoma hawaiense Edmonds, 1966] | | | `--- S. vastum (Selenka & Balow, 1883) [Sipunculus vastus Selenka & Balow, 1883; Sipunculus | | | {Phascolosomum} violaceus de Quatrefages, 1865; Siphonosoma crassum Spengel, 1919; | | | Sipunculus parvum Fischer, 1928; Siphonosoma vastum album (Shipley, 1899); Siphonosoma | | | vastum obscurum (Shipley, 1899)] | | |--o Siphonomecus Fischer, 1947 | | | `--- S. multicinctus Fischer, 1947 | | `--o Phascolopsis Fischer, 1950 [Golfingia {Phascolopsis} Fischer, 1950] | | `--- P. gouldii (Pourtales, 1851) [Sipunculus gouldii Pourtales, 1851] | `--o GOLFINGIIFORMES Cutler & Gibbs, 1985 | |--o Golfingiidae Stephen & Edmonds, 1972 | | |--o Golfingia Lankester, 1885 [Goldfingia {Dushana} Murina, 1975; Themiste {Centrosiphon} Shipley, 1903; | | | | Goldfingiella Stephen, 1964; Siphonoides Murina, 1967] | | | |--o {Spinata) Cutler, & Cutler, 1987 | | | | `--- G. {S.} pectinatoides (Cutler & Cutler, 1979) [Spinata pectinatoida Cutler & Cutler, 1979] | | | `--o {Golfingia} Lankester, 1885 | | | |--- G. {G.} anderssoni (Thiel, 1911) [Phascolosoma anderssoni Thiel, 1911] | | | |--- G. {G.} birsteini Murina, 1973 | | | |--- G. {G.} capensis (Teuscher, 1874) [Phascolosoma capensis Teuscher, 1874; Dendrostoma stephensoni | | | | Stephen, 1942; Themiste stephensoni Stephen, 1942] | | | |---o G. {G.} elongata (Keferstein, 1863) [Phascolosoma elongatum Keferstein, 1863; Phascolosoma | | | | cylindratum Keferstein, 1865; Phascolosoma obscurus de Quatrefages 1865; | | | | Phascolosoma obscurum Baird, 1868; Phascolosoma forbesii Baird, 1868; | | | | Phascolosoma oxyurum Baird, 1868; Phascolosoma tenuicinctum Baird, 1868; | | | | Phascolosoma delagei Hrubel, 1903; Phascolosoma charcoti Hrubel, 1906; | | | | Phascolosoma teres Hutton, 1903; Phascolosoma derjugini Gadd, 1911; | | | | Phascolosoma abyssorum punctata Hrubel, 1925; Phascolosoma cluthensis | | | | Stephen, 1931] | | | |--- G. {G.} iniqua (Sluiter, 1912) [Phascolosoma iniquum Sluiter, 1912; Phascolosoma rugosum | | | | Southern, 1913; Phascolosoma mutabile Southern, 1913] | | | |--o G. {G.} margaritacea (Sars, 1851) [Sipunculus margaritaceus Sars, 1951] | | | | |--- G. {G.} m. margaritacea (Sars, 1851) [Sipunculus margaritaceus Sars, 1951; Homalosoma laeve | | | | | Oersted, 1865; Phascolosoma oerstedii Keferstein, 1865; Phascolosoma oerstedii | | | | | Keferstein, 1865; Phascolosoma capsiforme Baird, 1868; Phascolosoma albidum Thiel, | | | | | 1875; Phascolosoma fulgens Thiel, 1875; Phascolosoma hanseni Danielssen & Koren, | | | | | 1881; Stephanostoma barentsii Horst, 1882; Phascolosoma antarcticum Michaelsen, | | | | | 1889; Phascolosoma fuscom Michaelsen, 1889; Phascolosoma georgianum Michaelsen, | | | | | 1889; Phascolosoma profundum Louis Roule, 1898; Phasacolosoma japonicum Ikeda, | | | | | 1904; Golfingia ikedai Fischer, 1950; Phascolosoma okinoseanum Ikeda, 1904; | | | | | Phascolosoma trybomi Thiel, 1905; Phasacolosoma socium Lanchester, 1908; | | | | | Phascolosoma nordenskjoldi Thiel, 1911; Phascolosoma meridionalis Gerould, 1913; | | | | | Phascolosoma glossipapillosa Satì, 1934; Phascolosoma noto Satì, 1934; | | | | | Phascolosoma stignum Satì, 1934; Phascolosoma soyo Satì, 1934; Golfingia | | | | | cantabriensis Edmonds, 1960] | | | | `--- G. {G.} m. ohlini (Thiel, 1911) [Phascolosoma ohlini Thiel, 1911; Phascolosoma pudicum | | | | Selenka, 1885; Phascolosoma mawsoni Benham, 1922; Goldfingia vulgaris antarctica | | | | (Murina, 1969)] | | | |--- G. {G.} mirabilis Murina, 1969 | | | |--- G. {G.} muricaudata (Southern, 1913) [Phascolosoma muricaudatum Southern, 1913; Phascolosoma | | | | appendiculatum Satì, 1934; Phascolosoma hudsonianum Chamberlain, 1920] | | | `--o G. {G.} vulgaris (de Blainville, 1827) [Sipunculus vulgaris de Blainville, 1827] | | | |--- G. {G.} v. vulgaris (de Blainville, 1827) [Sipunculus vulagris de Blainville, 1827; | | | | Phascolosoma astuta Selenka, 1885; Phascolosoma multipapillosa Hrubel, 1925; | | | | Phascolosoma selenkae Lanchester, 1905; Phascolosoma tropicum Sluiter, 1902; | | | | Goldfingia vulgaris visiculosus Murina, 1969; Phascolosoma longicolle Leuckart & | | | | Rueppel, 1828; Sipunculus papillosum Thompson, 1840; Syrinx harveyii Forbes, 1841; | | | | Sipunculus communis de Quatrefages, 1850; Sipunculus punctatissimus Gosse, 1853; | | | | Phascolosoma commune Keferstein, 1863; Sipunculus obscurus de Quatrefages, 1865; | | | | Phascolosoma luteum Thiel, 1875; Phascolosoma dubium Thiel, 1875; Phascolosoma | | | | validum Thiel, 1875; Goldfingia mackintoshii Lankester, 1885; Phascolosoma sanderi | | | | Collin, 1892; Phascolosoma owstoni Ikeda, 1904; Phascolosoma kolense Gadd, 1911; | | | | Phascolosoma solitarium Sluiter, 1912; Goldfingia {Dushana} adriatica Murina, | | | | 1975] | | | `--- G. {G.} v. herdmani (Shipley, 1903) [Centrosiphon herdmani Shipley, 1903; Goldfingia | | | queenslandensis Edmonds, 1956; Goldfingia liochros Cutler & Cutler, 1979] | | |--o Nephasoma Pergament, 1940 [Goldfingia {Phascoloides} Fischer, 1950] | | | |--o {Cutlerensis) Popkov, 1991 | | | | `--- N. {C.} rutilofuscum (Fischer, 1916) [Aspidosiphon rutilofuscus Fischer, 1916] | | | `--o {Nephasoma} Pergament, 1940 | | | |--o N. {N.} abyssorum (Koren & Danielsen, 1875) [Phascolosoma abyssorum Koren & Danielsen, 1875] | | | | |--- N. {N.} a. abyssorum (Koren & Danielsen, 1875) [Phascolosoma abyssorum Koren & Danielsen, | | | | | 1875; Phascolosoma incompositum Sluiter, 1912] | | | | `--- N. {N.} a. benhami (Stephen, 1948) [Phascolosoma benhami Stephen, 1948] | | | |--- N. {N.} bulbosum (Southern, 1913) [Phascolosoma bulbosum Southern, 1913] | | | |--- N. {N.} capilleforme (Murina, 1973) [Goldfingia capilleformis Murina, 1973; Goldfingia glacialis | | | | Murina, 1964] | | | |--- N. {N.} confusum (Sluiter, 1902) [Phascolosoma confusum Sluiter, 1902; Goldfingia confusa | | | | zarenkovi Murina, 1974] | | | |--- N. {N.} constricticervix (Cutler, 1969) [Goldfingia constricicervix Cutler, 1969] | | | |--- N. {N.} constrictum (Souther, 1913) [Phascolosoma constrictum Souther, 1913] | | | |--- N. {N.} cutleri (Murina, 1975) [Goldfingia cutleri Murina, 1975] | | | |--o N. {N.} diaphanes (Gerould, 1913) [Phascolosoma diaphanes Gerould, 1913] | | | | |--- N. {N.} d. diaphanes (Gerould, 1913) [Phascolosoma diaphanes Gerould, 1913; Phascolosoma | | | | | minutum Thiel, 1911; Phascolosoma anceps Sluiter, 1912; Phascolosoma sabellariae | | | | | Gerould, 1913; Phascolosoma improvisum Gerould, 1913; Phascolosoma cinctum | | | | | Gerould, 1913; Goldfingia sectile Murina, 1974] | | | | `--- N. {N.} d. corrugatum Cutler & Cutler, 1986 [Goldfingia schuttei Murina, 1964] | | | |--- N. {N.} eremita (Sars, 1851) [Sipunculus eremita Sars, 1851; Sipunculus borealis de Quatrefages, | | | | 1865; Phascolosoma digitatum Thiel, 1875; Goldfingia elachea Fischer, 1952] | | | |--- N. {N.} filiforme (Sluiter, 1902) [Phascolosoma filiforme Sluiter, 1902] | | | |--- N. {N.} flagriferum (Selenka, 1885) [Phascolosoma flagriferum Selenka, 1885] | | | |--- N. {N.} laetmophilum (Fischer, 1952) [Goldfingia laetmophila Fischer, 1952] | | | |--- N. {N.} lilljeborgi (Danielssen & Koren, 1180) [Phascolosoma lilljeborgii Danielssen & Koren, | | | | 1880; Nephasoma marinki Pergament, 1940; Onchnesoma glaciale Danielssen & | | | | Koren, 1880] | | | |--- N. {N.} minutum (Keferstein, 1862) [Phascolosoma minutum Keferstein, 1862; Phascolosoma | | | | sabellariae Thiel, 1905; Phascolosoma improvisum Thiel, 1905; Phascolosoma | | | | anceps Thiel, 1905] | | | |--- N. {N.} multiaraneusa (Murina, 1967) [Goldfingia multiaraneusa Murina, 1967] | | | |--- N. {N.} novaezealandiae (Benham, 1904) [Phascolosoma novae-zealandiae Benham, 1904] | | | |--o N. {N.} pellucidum (Keferstein, 1865) [Phascolosoma pellucidum Keferstein, 1865] | | | | |--- N. {N.} p. pellucidum (Keferstein, 1865) [Phascolosoma pellucidum Keferstein, 1865; | | | | | Phascolosoma riisei Keferstein, 1865; Phascolosoma cinereum Gerould, 1913; | | | | | Phascolosoma verrillii Gerould, 1908; Phascolosoma sluiteri ten Broeke, 1925; | | | | | Goldfingia coriacea Fischer, 1950; Goldfingia eremita australe Wesenberg-Lund, | | | | | 1959] | | | | `--- N. {N.} p. subhamatum (Sluiter, 1902) [Phascolosoma subhamatum Sluiter, 1902] | | | |--- N. {N.} rimicola (Gibbs, 1973) [Goldfingia rimicola Gibbs, 1973] | | | |--- N. {N.} schuettei (Augener, 1903) [Phascolosoma schüttei Augnener, 1903] | | | |--- N. {N.} tasmaniense (Murina, 1964) [Goldfingia tasmaniensis Murina, 1964] | | | |--- N. {N.} vitjazi (Murina, 1964) [Goldfingia vitjazi Murina, 1964] | | | `--o N. {N.} wodjanizkii (Murina, 1973) [Goldfingia wodjanizkii Murina, 1973] | | | |--- N. {N.} w. wodjanizkii (Murina, 1973) [Goldfingia wodjanizkii Murina, 1973] | | | `--- N. {N.} w. elisae (Murina, 1977) [Goldfingia elisae Murina, 1977] | | `--o Thysanocardia Fischer, 1950 [Golfingia {Thysanocardia} Fischer, 1950] | | |--- T. catharinae (Grübe, 1868) [Phascolosoma catharinae Grübe, 1868; Phascolosoma semperi Selenka & de | | | Man, 1883; Phascolosoma martensi Collin, 1901; Phascolosoma procera Gerould, 1913] | | |--- T. nigra (Ikeda, 1904) [Phascolosoma nigrum Ikeda, 1904; Phascolosoma pavlenkoi Ostroumov, 1909; | | | Phascolosoma hyugensis Sati, 1934; Phascolosoma zenibakense Ikeda, 1924; Phascolosoma | | | onagawa Sati, 1937; Phascolosoma hozawai Sati, 1937; Goldfingia pugettensis Fischer, 1952; | | | Goldfingia macginitiei Fischer, 1952; Goldfingia catharinae Cutler, 1977] | | `--- T. procera (Mibius, 1875) [Phascolosoma procerum Mibius, 1875] | |--o Phascolionidae Cutler & Gibbs, 1985 [Phascoliidae Cutler & Gibbs, 1985] | | |--o Phascolion Thiel, 1875 | | | |--o {Isomya} Cutler & Cutler, 1985 | | | | |--- P. {I.} convestitum (Sluiter, 1902) [Phascolion convestitus Sluiter, 1902; Phascolion | | | | | mediterraneum Fischer, 1922; Phascolion beklemischevi Murina, 1964] | | | | |--- P. {I.} gerardi Rice, 1993 | | | | |--- P. (I.} hedraeum Selenka & de Man, 1883 [Phascolion dentalicola Sati, 1937; Phascolion | | | | | kurchatovi Murina, 1947] | | | | |--- P. {I.} lucifugax Selenka & de Man, 1883 | | | | |--- P. {I.} microsperhoidis (Cutler & Duffy, 1972) [Phascolion microsheroidi Cutler & Duffy, 1972] | | | | `--- P. {I.} tuberculosum Thiel, 1875 [Tylosoma lytkenii Koren & Daneilssen, 1875; Phascolion | | | | pallidum Koren & Danielssen, 1875; Phascolion hirondellei Sluiter, 1900; Phascolion | | | | temporariae Edmonds, 1976] | | | |--o {Lesenka} Gibbs, 1985 | | | | |--- P. {L.} collare Selenka & de Man, 1883 | | | | |--- P. {L.} cryptum (Hendrix, 1975) [Phascolion cryptus Hendrix, 1975] | | | | |--- P. {L.} hupferi Fischer, 1895 [Phascolion indicus Murina, 1974] | | | | |--- P. {L.} rectum Ikeda, 1904 | | | | `--o P. {L.} valdiviae Fischer, 1916 | | | | |--- P. {L.} v. valdiviae Fischer, 1916 | | | | `--- P. {L.} v. sumatrense Fischer, 1916 [Phascolion murrayi Stephen, 1941] | | | |--o {Montuga} Gibss, 1985 | | | | `--o P. {M.} lutense Selenka, 1885 | | | | |--- P. {M.} l. lutense Selenka, 1885 [Phascolion canum Cutler & Cutler, 1980] | | | | `--- P. {M.} l. pacificum Murina, 1957 | | | |--o {Phascolion} Thiel, 1875 | | | | |--- P. {P.} abnorme Fischer, 1895 [Phascolion heteropapillosum Wesenberg-Lund, 1963] | | | | |--- P. {P.} bogorovi Murina, 1973 | | | | |--- P. {P.} caupo Hendrix, 1975 | | | | |--- P. {P.} hibridum Murina, 1981 | | | | |--- P. {P.} medusae Cutler & Cutler, 1980 | | | | |--- P. {P.} megaethi Cutler & Cutler, 1979 | | | | |--- P. {P.} pharetratum Sluiter, 1902 | | | | |--- P. {P.} psammophilus Rice, 1993 | | | | |--- P. {P.} robertsoni Stephen & Robertson, 1952 | | | | |--o P. {P.} strombus (Montagu, 1804) [Sipunculus strombus Montagu, 1804] | | | | | |--- P. {P.} s. strombus (Montagu, 1804) [Sipunculus strombus Montagu, 1804; Sipunculus dentalii | | | | | | Gray, 1828; Sipunculus bernnhardus Forbes, 1841; Sipunculus capitatus Rathke, 1843; | | | | | | Sipunculus caementarius de Quatrefages, 1865; Phascolosoma hamulatum Packard, 1867; | | | | | | Phascolion spetsbergense Thiel, 1875; Phascolosoma tubicola Verrill, 1873; Sipunculus | | | | | | concharum Oersted, 1844; Phascolion alberti Sluiter, 1900; Phascolion artificiosus | | | | | | Ikeda, 1904; Phascolion mogadrense Sluiter, 1912; Phascolosoma intermedium Southern, | | | | | | 1913; Phascolion africanum Fischer, 1923; Phascolion brotzkajae Murina, 1964; | | | | | | Phascolion tortum Edmonds, 1976; Phascolion anomalus Murina, 1981] | | | | | `--- P. {P.} s. cronullae Edmonds, 1980 | | | | `--- P. {P.} ushakovi Murina, 1975 | | | `--o {Villiophora} Cutler & Cutler, 1985 | | | `--- P. {V.} cirratum (Murina, 1968) [Phascolion cirratus Murina, 1968] | | `--o Onchnesoma Koren & Danielssen, 1875 | | |--- O. intermedium Murina, 1976 | | |--- O. magnibathum Cutler, 1969 | | |--o O. squamatum (Koren & Danielsen, 1875) [Phascolosoma squamatum Koren & Danielssen, 1875] | | | |--- O. s. squamatum Murina, 1976 (Koren & Danielsen, 1875) [Phascolosoma squamatum Koren & | | | | Danielssen, 1875] | | | `-- O. s. oligopapillosum Cutler, 1984 | | `--o O. steenstrupii Koren & Danielssen, 1875 | | |--- O. s. steenstrupii Koren & Danielssen, 1875 [Phascolion dogieli Murina, 1964] | | `--- O. s. nudum Cutler & Cutler, 1985 | `--o Themsitidae Cutler & Gibbs, 1985 | `--o Themiste Gray, 1828 [Dendrostomum Grübe & Oersted, 1858; Dendrostoma Keferstein, 1865] | |--o {Themiste} Gray, 1828 | | |--- T. {T.} alutacea (Grübe & Oersted, 1858) [Dendrostomum alutaceum Grübe & Oersted, 1858; | | | Sipunculus orbiniensis de Quatrefages, 1865; Dendrostomum petrolicum Amor, 1964; | | | Dendrostomum rosaceum Amor, 1964] | | |--- T. {T.} blanda (Selenka & Man, 1883) [Dendrostomum blandum Selenka & Man, 1883] | | |--- T. {T.} dyscrita (Fischer, 1952) [Dendrostomum dyscritum Fischer, 1952] | | |--- T. {T.} hennahi Gray, 1828 [Sipunculus rapa de Quatrefages, 1865; Dendrostomum ramonum de | | | Quatrefages, 1865; Dendrostoma peruvianum Collin, 1892; Dendrostoma mytheca Chamberlin, | | | 1920; Dendrostoma zostericolum Chamberlin, 1920; Dendrostoma perimeces Fischer, 1928; | | | Dendrostomum lissum Fischer, 1952; Dendrostomum schmitti Fischer, 1952] | | `--- T. {T.} pyroides (Chamberlin, 1920) [Dendrostoma pyroides Chamberlin, 1920; Dendrostoma petraeum | | Fischer, 1928; Dendrostoma hexadactylum Sati, 1930] | `--o {Lagenopsis} Edmonds, 1980 | |--- T. {L.} cymodoceae (Edmonds, 1956) [Dendrostoma cymodoceae Edmonds, 1956] | |--- T. {L.} dehamata (Kesteven, 1903) [Dendrostoma dehamatum Kesteven, 1903; Dendrostoma ellipticum | | Sati, 1934; Dendrostoma fischeri Amor, 1964] | |--- T. {L.} laginiformis Baird, 1868 [Dendrostoma cymodoceae Edmonds, 1956] | |--- T. {L.} cymodoceae (Edmonds, 1956) [Dendrostoma signifer Selenka & de Man, 1883; Phascolosoma | | pyriformis Lanchester, 1905; Dendrostoma tropicum Sati, 1942; Dendrostoma stephensoni | | Stephen, 1942; Dendrostoma minor Chin, 1947; Dendrostoma robertsoni Stephen & Roberston, | | 1952] | |--o T. {L.} minor (Ikeda, 1904) [Dendrostoma minor Ikeda, 1904] | | |--- T. {L.} m. minor (Ikeda, 1904) [Dendrostoma minor Ikeda, 1904; Dendrostomum fuscum Edmonds, | | | 1960; Golfingia coriacea Murina, 1972] | | `--- T. {L.} m. huttoni (Benham, 1903) [Phascolosoma huttoni Benham, 1903] | `--- T. {L.} variospina Edmonds, 1956 `--o PHASCOLOSOMATIDEA Cutler & Gibbs, 1985 [PHASCOLOSOMIDA Cutler & Gibbs, 1985] |--o PHASCOLOSOMATIFORMES Cutler & Gibbs, 1985 [PHASCOLOSOMAFORMES Cutler & Gibbs, 1985] | `--o Phascolosomatidae Stephen & Edmonds, 1972 | |--o Phascolosoma Leuckart, 1828 [Phascolosomum Diesing, 1851; Phymosomum de Quatrefages, 1865; Phymosoma | | | Selenka, 1883; Physcosoma Selenka, 1897] | | |--o {Fischerana} Stephen, 1964 | | | |--- P. {F.} capitatum (Gerould, 1913) [Physcosoma capitatum Gerould, 1913; Physcosoma abyssorum | | | | Southern, 1913; Goldfingia {Fisherana} savalovi Murina, 1974] | | | `--- P. {F.} lobostomum Fischer, 1895 [Golfingia {Apionsoma} lobostomum Cutler, 1979; Fischerana | | | wasini Lanchester, 1905] | | `--o {Phascolosoma} Leuckart, 1828 | | |--o P. {P.} agassizii Keferstein, 1866 | | | |--- P. {P.} a. agassizii Keferstein, 1866 [Phascolosoma lordi Baird, 1868; Phascolosoma japonicum | | | | Grübe, 1877; Physcosoma glaucum Sati, 1930; Physcosoma yezoense Ikeda, 1924; | | | | Physcosoma formosense Sati, 1939; Phascolosoma {Rueppellisoma} golikovi Murina, | | | | 1975] | | | `--- P. {P.} a. kurilense (Sati, 1937) [Physcosoma kurilense Sati, 1937] | | |--- P. {P.} albolineatum Baird, 1868 [Phymosoma microdonton Sluiter, 1886; Phascolosoma | | | multiannulata Wesenberg-Lund, 1954; Phascolosoma andamanensis Johnson, 1971] | | |--- P. {P.} annulatum Hutton, 1879 [Phascolosoma scolops mossambiciense Augener, 1903; Phascolosoma | | | scolops tasmaniense Fischer, 1914] | | |--- P. {P.} arcuatum (Gray, 1828) [Siphunculus arcuatus Gray, 1828; Phymosoma lurco Selenka & de | | | Man, 1883; Phascolosoma hizophora Sluiter, 1891; Physcosoma ambonense Fischer, 1896; | | | Phymosoma deani Ikeda, 1905; Physcosoma esculenta Chen & Yeh, 1958] | | |--o P. {P.} glabrum (Sluiter, 1902) [Physcosoma glabrum Sluiter, 1902] | | | |--- P. {P.} g. glabrum (Sluiter, 1902) [Physcosoma glabrum Sluiter, 1902; Phymosoma microdontodon | | | | Shipley, 1898; Physcosoma funafutiense Fischer, 1914] | | | `--- P. {P.} g. multiannulatum Wesenberg-Lund, 1954 | | |--- P. {P.} granulatum Leuckart, 1828 [Sipunculus flavus Risso, 1826; Sipunculus tigrinus Risso, | | | 1826; Sipunculus genuensis de Blainville, 1827; Phascolosoma laeve Cuvier, 1830; | | | Sipunculus verrucosus Cuvier, 1830; Sipunculus levis Cuvier, 1830; Sipunculus papillosum | | | Thompson, 1840; Sipunculus multitorquatus de Quatrefages, 1865; Phascolosoma fasciatum | | | Baird, 1868; Phascolosoma jeffreysii Baird, 1868; Phascolosoma loveni Koren & | | | Danielssen, 1877; Physcosoma herowardii Hrubel, 1903; Physcosoma lanzarotae Harms, 1921] | | |--- P. {P.} maculatum (Sluiter, 1886) [Phymosoma maculatum Sluiter, 1886] | | |--- P. {P.} meteori (Hrubel, 1904) [Phymosoma meteori Hrubel, 1904] | | |--- P. {P.} nigrescens Keferstein, 1865 [Phascolosoma puntarenae Grübe & Oersted, 1858; Sipunculus | | | {Phymosomum} plicatus de Quatrefages, 1865; Phascolosoma varians Keferstein, 1865; | | | Phascolosoma agassizii Keferstein, 1867; Phascolosoma plainispinosum Baird, 1868; | | | Phymososma spengeli Sluiter, 1886; Phymososoma duplicigranulatum Sluiter, 1886; | | | Phymosoma lacteum Sluiter, 1886; Phymosoma diaphanes Sluiter, 1886; Physcosma extortum | | | Sluiter, 1902; Physcosoma evisceratum Lanchester, 1905; Physcosoma minutum ten Broeke, | | | 1925; Physcosoma horsti ten Broeke, 1925] | | |--- P. {P.} noduliferum Stimpson, 1855 [Sipunculus tuberculatus Gray, 1828; Sipunculus {Phymosomum} | | | nodulosus de Quaterfages, 1865; Sipunculus {Phymosomum} javenensis de Quatrefages, 1865; | | | Phascolosoma grayi Baird, 1868] | | |--- P. {P.} pacificum Keferstein, 1866 [Sipunculus (Phymosomum} javenensis de Quatrefages, 1865; | | | Phasacolosoma asperum Grübe, 1868] | | |--- P. {P.} perlucens Baird, 1868 [Sipunculus {Phascolosomum} vermiculus de Quatrefages, 1865; | | | Phymosoma dentigerum Selenka & de Man, 1883; Physcosoma thomense Augener, 1903; | | | Aspidosiphon insularis Lanchester, 1905; Physcosoma microdentigerum ten Broeke, 1925; | | | Phascolosoma spinosum Johnson, 1971] | | |--- P. {P.} saprophagicum Gibbs, 1987 | | |--- P. {P.} scolops (Selenka & de Man, 1883) [Phymosoma scolops Selenka & de Man, 1883; Phymosoma | | | scolops mossambiciense Selenka & de Man, 1883; Phascolosoma carneum Leuckart & Rüppell, | | | 1828; Sipunculus {Phymosomum} guttatus de Quatrefahges, 1865; Phymosoma psaron Sluiter, | | | 1886; Phymosoma spongicola Sluiter, 1902; Phymosoma nahaense Ikeda, 1904; Phymosoma | | | socium Lanchester, 1905; Phascolosoma rottnesti Edmonds, 1956; Phascolosoma dunwichi | | | Edmonds, 1956; Phascolosoma riukiuensis Murina, 1975] | | |--- P. {P.} stephensoni (Stephen, 1942) [Physcosoma stephensoni Stephen, 1942; Phascolosoma laeve | | | Keferstein, 1862; Sipunculus {Phymosomum} spinicauda de Quatrefages, 1865; Physcosoma | | | lanzarotae Harms, 1921; Phascolosoma heronis Edmonds, 1956; Phascolosoma granulatum | | | Murina, 1981] | | `--- P. {P.} turnerae Rice, 1985 [Phascolosoma kapalum Edmonds, 1985] | |--o Antillesoma Stephen & Edmonds, 1972 [Phascolosoma {Antillesoma} Stephen & Edmonds, 1972] | | `--- A. antillarum (Grübe & Oersted, 1858) [Phascolosoma antillarum Grübe & Oersted, 1858; Phascolosoma | | fuscum Keferstein, 1862; Sipunculus {Aedematosomum} glans de Quatrefages, 1865; | | Phascolosoma nigriceps Baird, 1868; Phymosoma asser Selenka & de Man, 1883; Phymosoma | | pelam Selenka & de Man, 1883; Physcosoma weldoni Shipley, 1892; Phymosoma | | onomichianum Ikeda, 1904; Physcosoma gaudens Lanchester, 1905; Physcosoma similkis | | Chen & Yeh, 1958; Golfingia mokyevskii Murina, 1964; Phascolosoma schmidti Murina, | | 1975] | `--o Apionsoma Sluiter, 1902 [Golfingia {Mitosiphon} Fischer, 1950; Golfingia {Phascolana} Wesenberg-Lund, | | 1959] | |--o Apionsoma Sluiter, 1902 | | |?—- A. {A.} immunitum (Sluiter, 1902) [Phascolsosoma immunitum Sluiter 1902][sp. inquirenda] | | |?—- A. {A.} papilliferum (Keferstein, 1865) [Phascolsosoma papilliferum Keferstein, 1865][sp. | | | inquirenda] | | |?—- A. {A.} reconditum (Sluiter, 1900) [Phascolsosoma reconditum Sluiter 1900][sp. inquirenda] | | |--- A. {A.} misakianum (Ikeda, 1904) [Phascolosoma misakianum Ikeda, 1904; Phascolosoma hespera | | | Chamberlin, 1920; Golfingia {Phascolana} longirostris Wesenberg-Lund, 1959; Golfingia | | | {Phasacolana} tenuissima Wesenberg-Lund, 1959] | | |---o A. {A.} murinae {Cutler, 1969) [Golfingia {Mitosiphon} murinae Cutler, 1969] | | | |--- A. {A.} m. murinae (Cutler, 1969) [Golfingia murinae Cutler, 1969; Golfingia hespera Murina, | | | | 1964] | | | `--- A. {A.} m. bilobatae (Cutler, 1969) [Golfingia {Mitosiphon} murinae bilobatae Cutler, 1969] | | `--- A. {A.} trichoscephalus Luiter, 1902 [Phascolosoma pusillum Sluiter, 1912] | `--o Edmondsius Gibbs & Cutler, 1987 [Phascolosoma {Satinus} Stpehen & Edmonds, 1972] | `--- A. {E.} pectinatum (Keferstein, 1867) [Phascolosoma pectinatum Keferstein, 1867; Phascolosoma | abnorme Sluiter, 1886] `--o ASPIDOSIPHONIFORMES Cutler & Gibbs, 1985 `--o Aspidosiphonidae Baird, 1868 |--o Aspidosiphon Diesing, 1851 [Pseudoaspidosiphon Baird, 1868] | |--o {Akrikos} Cutler & Cutler, 1989 | | |--- A. {A.} albus Murina, 1967 [Akrikos hartmeyeri Wesenberg-Lund, 1957] | | |--- A. {A.} mexicanus (Murina, 1967) [Golfingia mexicana Murina, 1967; Aspidosiphon longirhynchus | | | Cutler & Cutler, 1980] | | |--- A. {A.} thomassini Cutler & Cutler, 1979 | | |--- A. {A.} venabulum Selenka & de Man, 1883 | | `--- A. {A.} zinni Cutler, 1969 | |--o {Aspidosiphon} Diesing, 1851 | | |--- A. {A.} elegans (Chamisso & Eysenhardt, 1821) [Sternaspis elegans Chamisso & Eysenhardt, 1821; | | | Aspidosiphon ravus Sluiter, 1886; Aspidosiphon exilis Sluiter, 1886; Aspidosiphon | | | spinosus Sluiter, 1902; Aspidosiphon brocki Augener, 1903; Aspidosiphon spinalis Ikeda, | | | 1904; Aspidosiphon carolinensis Sati, 1935; Aspidosiphon homoyarium Johnson, 1964] | | |--- A. {A.} exiguus Edmonds, 1974 | | |--- A. {A.} gosnoldi Cutler, 1981 [Aspidosiphon spinalis Cutler, 1973] | | |--o A. {A.} gracilis (Baird, 1868) [Pseudaspidosiphon gracile Baird, 1868] | | | |--- A. {A.} g. gracilis (Baird, 1868) [Pseudaspidosiphon gracile Baird, 1868] | | | `--- A. {A.} g. schnehageni Fischer, 1913 | | |--- A. {A.} misakiensis Ikeda, 1904 [Aspidosiphon hartmeyeri Fischer, 1919; Aspidosiphon gerouldi | | | ten Broeke, 1925; Aspidosiphon speculatus Saiz, 1886] | | |--- A. {A.} muelleri Diesing, 1851 [Sipunculus scutatus Müller, 1844; Aspidosiphon clavatus Diesing, | | | 1851; Sipunculus cochlearius Valenciennes, 1854; Lesinia farcimen Schmidt, 1854; | | | Aspidosiphon eremita Diesing, 1859; Phascolosoma radiata Alder, 1860; Sipunculus | | | heterocyathi McDonald, 1862; Aspidosiphon jukesii Baird, 1873; Aspidosiphon mirabilis | | | Thiel, 1875; Aspidosiphon armatum Danielssen & Koren, 1880; Aspidosiphon tortus Selenka | | | & Bülow, 1883; Aspidosiphon heteropsammiarum Bouvier, 1894; Aspidosiphon michelini | | | Bouvier, 1894; Aspidosiphon corallicola Sluiter, 1902; Aspidosiphon imbellis Sluiter, | | | 1902; Aspidosiphon inquilinus Sluiter, 1902; Aspidosiphon exhaustum Sluiter, 1912; | | | Aspidosiphon pygmaeus Fischer, 1921; Aspidosiphon kovaleskii Murina, 1964; Aspidosiphon | | | hispitrofus LiGreci, 1980] | | `--- A. {A.} spiralis Sluiter, 1902 | `--o {Paraspidosiphon} Stephen, 1964 | |--- A. {P.} coyi de Quatrefages, 1865 [Phascolosoma truncatum Keferstein, 1867] | |--- A. {P.} fischeri ten Broeke, 1925 [Aspidosiphon fischerti cubanus Murina, 1967] | |--- A. {P.} laevis (de Quatrefages, 1865) [Aspidosiphon laeve de Quaterfages, 1865; Aspidosiphon | | cumingii Baird, 1868; Aspidosiphon major Vaillant, 1871; Aspidosiphon klunzingeri | | Selenka & Bülow, 1883; Aspidosiphon gigas Sluiter, 1884; Aspidosiphon anuglatus Ikeda, | | 1904; Aspidosiphon speciosus Gerould, 1913; Aspidosiphon grandis Sati, 1939; | | Aspidosiphon pachydermatus Wesenberg-Lund, 1937; Aspidosiphon brasiliensis Cordero & | | Mello-Leitao, 1952; Aspidosiphon johnstoni Edmondsd, 1980; Aspidosiphon quatrefagei | | Saiz, 1984] | |--- A. {P.} parvulus Gerould, 1913 [Aspidosiphon spinoso-scutatus Fischer, 1922] | |--- A. {P.} planoscutatus Murina, 1968 | |--- A. {P.} steenstrupii Diesing, 1859 [Aspidosiphon fuscus Sluiter, 1881; Aspidosiphon speculator | | Selenka, 1885; Aspidosiphon semperi ten Broeke, 1925; Aspidosiphon makoensis Sati, 1939; | | Aspidosiphon trinidensis Cordero & Mello-Leitao, 1952; Aspidosiphon exostomum Johnson, | | 1964; Aspidosiphon ochrus Cutler & Cutler, 1979] | `--- A. {P.} tenuis Sluiter, 1886 [Aspidosiphon levis Sluiter, 1886; Aspidosiphon ambonensis Augener, | 1903; Aspidosiphon formosanus Sati, 1939; Aspidosiphon havelockensis Haldar, 1978] |--o Lithacrosiphon Shipley, 1902 | |--o L. cristatus (Sluiter, 1902) [Aspidosiphon cristatus Sluiter, 1902] | | |--- L. c. cristatus (Sluiter, 1902 [Aspidosiphon cristatus Sluiter, 1902; Aspidosiphon uniscutatus | | | Ikeda, 1904; Lithacrosiphon kukenthali Fischer, 1922; Lithacrosiphon indicus | | | Fischer, 1922; Lithacrosiphon odhneri Fischer, 1922; Lithacrosiphon alticornis | | | ten Broeke, 1925; Lithacrosiphon portidis ten Broeke, 1925; Lithacrosiphon | | | gurjanovae Murina, 1967] | | `--- L. c. lakshadweepensis Haldar, 1991 | `--- L. maldivensis Shipley, 1902 `--o Cloeosiphon Grübe, 1868 [Loxosiphon de Quatrfages, 1865] `--- C. aspergillus (de Quatrefages, 1865) [Loxosiphon aspergillus de Quatrefages, 1865; Cloeosiphon mollis Selenka & Bülow, 1883; Cloeosiphon javanicum Sluiter, 1886; Cloesiphon japonicum Ikeda, 1904; Cloeosiphon carolinum Ikeda, 1904] http://www.mcz.harvard.edu/Departments/InvertZoo/as.fldr/cutler/siptaxa.htm