After Ebach & Ahyong, 2001, and Adrain, 2003
<==o †Lichidae Hawle & Corda, 1847 |-- †Lichinae Hawle & Corda, 1847 `--o †Trochurinae Phleger, 1936 |--o †Dicranogmus Hawle & Corda, 1847 | |-- †D. pustulatus Hawle & Corda, 1847 | |-- †D. aequalis (Törnquist, 1884) [Lichas aequalis Törnquist, 1884] | |-- †D. gregalis Chugaeva, 1983 | |-- †D. guizhouensis Wu, 1977 | |-- †D. scabrosus Zhou & Zhou, 1982 | |-- †D. skinneri Perry & Chatterton, 1977 | `-- †D. wynni Adrain, 2003 |--o †Hemiarges Gürich, 1901 | |-- †H. wesenbergensis | `-- †H. turneri `--+-- †Radiolichas Reed, 1923 |-- †Craspedarges Gürich, 1901 |-- †Ceratarges Gürich, 1901 |-- †Eifliarges, Richter & Richter, 1917 |-- †Mephiarges Richter & Richter, 1930 |-- †Terranovia Maksimova, 1977 |-- †Richterarges Phleger, 1936 |--o †Borealarges Adrain, 1994 | |?- †B. simplex (Barrande, 1846) [†Lichas simplex Barrande, 1846; †Dicranogmus simplex (Barrande, 1846)] | |?- †B. convexus (Tomczykowa, 1991) [Richterarges convexus Tomczykowa, 1991] | |?- †B. bucklandii (Milne Edwards, 1840) | |?- †B. kielcensis (Tomczykowa, 1993) [Richterarges kielcensis Tomczykowa, 1993] | |--+-- †B. warholi Adrain, 2003 | | `--+-- †B. tuckerae Adrain, 1994 | | `--+-- †B. nicoae Adrain, 2003 | | `-- †B. yulei Adrain, 2003 | `--+-- †B. calei Adrain, 1994 | `--+-- †B. morrisoni Adrain, 1994 | `--+-- †B. reedi Adrain, 1994 | `-- †B. mikulicorum Perry & Chatterton, 1977 `--o †Acanthopyge Hawle & Corda, 1847 |--o †(Acanthopyge) Hawle & Corda, 1847 | |-- †A. (A.) duplicispinata Kaneko, 1984 | |-- †A. (A.) granulosa (Roemer, 1852) | |-- †A. (A.) haueri Barrande, 1846 [A. (A.) leuchtenbergii] | `-- †A. (A.) mediasulcatus (Kaneko, 1984) |--o †(Jasperia) Thomas & Holloway, 1988 | `-- †A. (J.) bifida (Edgell, 1955) |--o †(Pernuaspis) Pribyl, 1949 [Lobopyge (Nitidulopyge)] | |-- †A. (P.) helga (Pribyl et al., 1986) | |-- †A. (P.) longispina Pribyl, 1949 | |-- †A. (P.) minuta (Barrande, 1846) | `-- †A. (P.) sexlobata (Roemer, 1855) `--o †(Lobopyge) Pribyl & Erben, 1952 [Benelopyge] sensu Ebach & Ahyong, 2001 |-- †A.? (L.) parapleura Curtis & Lane, 1998 |-- †A. (L.) altirhachis Chernysheva, 1951 |-- †A. (L.) contusa (Hall & Clarke, 1888) |-- †A. (L.) hexapteryx Pribyl & Erben, 1952 |-- †A. (L.) longiaxis (Maksimova, 1968) |-- †A. (L.) niobe (Basse, 1998) |-- †A. (L.) parvulus (Novak, 1890) |-- †A. (L.) permarginata (Pribyl & Erben, 1952) |-- †A. (L.) pervasta (Pribyl et al., 1986 |-- †A. (L.) pulex (Haas, 1968) |-- †A. (L.) pusilla (Angelin, 1854) |-- †A. (L.) pustulosa (Morzadec, 1983) |-- †A. (L.) sibirica (Chernysheva, 1951) |-- †A. (L.) sinuata (Ratte, 1886) |-- †A. (L.) rohri `--+-- †A. (L.) pustulosa `--+--+-- †A. (L.) edgecombei Ebach, 2001 | `-- †A. (L.) erinacea (Haas, 1968) `--+--+-- †A. (L.) sinuata (Ratte, 1886) | `--+-- †A. (L.) australis (McCoy, 1876) | `--+-- †A. (L.) australiformis Chatterton, Johnston & Campbell, 1979 | `-- †A. (L.) campbelli Chatterton & Wright, 1986 `--+-- †A. (L.) consanguinea (Clarke, 1894) `--+--+-- †A. (L.) ballivaini (Kozlowski, 1923) [Lichas; Benelopyge] | `--+-- †A. (L.) richteri (Vanek, 1959) | `-- †A. (L.) uralensis (Maksimova, 1979) `--+-- †A. (L.) brankiensis (Barrande, 1872) `--+-- †A. (L.) decheni (Holzapfel, 1895) `--+--+-- †A. (L.) parva (Barrande, 1846) | `-- †A. (L.) pragensis (Boucek, 1933) `--+--+-- †A. (L.) brevis (Maksimova, 1968) | `-- †A. (L.) limbata (Maksimova, 1968) `--+-- †A. (L.) sp. [Victoria, Australia] [Holloway & Neil, 1982 (from Ebach & Ahyong, 2001)] `--+--+-- †A. (L.) orientalis (Wu, 1977) | `-- †A. (L.) parapleura Curtis & Lane, 1998 `--+-- †A. (L.) docekali (Venek, 1959) `--+-- †A. (L.) hirsuta (Fletcher, 1850) `-- †A. (L.) trinodis (Ellermann, 1992)
Reference(s):
- Adrain, J. M., 2003: Validity and composition of the Silurian trilobite genera Borealarges and Dicranogmus, wit new species from the Canadian Arctic.
–Canadian Journal of Earth Sciencies: Vol. 40, pp. 749-763 - Ebach, M. C. & Ahyong, S. T., 2001: Phylogeny of the trilobite subgenus Acanthopyge (Lobopyge).
–Cladistics: Vol. 17, #1, pp. 1-10 - Gon III, S. M., 2002: Systematic listing of Trilobite families.
–iNet: http://www.aloha.net/~smgon/trisystem.htm