Mikko's Phylogeny Archive Main Groups of Protostomes   Scorpionida - scorpions

After Størmer, Petrunkevitch & Hedgpeth, 1955 and Stockwell, 1996 ["The Tree of Life"]

<==o Scorpionida Latreille, 1817 (scorpions; skorpionit)
   |--o †Protoscorpionina Petrunkevitch, 1949 [pro Protoscorpiones Petrunkevitch, 1949]
   |  |-- †Allopalaeophonus Kjellesvig-Waering, 1986 [Allopalaeophonidae]
   |  |-- †Dolichophonus loudonensis (Laurie, 1889) Petrunkevitch, 1953 [Palaeophonus loudonensis Laurie, 1889] [Dolichophonidae Petrunkevitch, 1953]
   |  |-- †Palaeoscorpius devonicus Lehmann, 1944 [Palaeoscorpiidae Lehmann, 1944; Palaeoscorpionoidea Lehmann, 1944]
   |  `--o †Palaeophonus Thorell & Lindström, 1885 [Palaeophoneus Thorell & Lindström, 1884, lapsus calami; Palaeophonidae Thorell & Lindström, 1885; Palaeophonoidea Thorell & Lindström, 1885]]
   |     |-- †P. nuncius Thorell & Lindström, 1885
   |     `-- †P. caledonicus Hunter,
   `--+--o †Palaeoscorpiones Lehmann, 1944
      |  |--o †Archaeoctonidae Petrunkevitch, 1949 [Archaeoctonoidea Petrunkevitch, 1949]
      |  |  |-- †Archaeoctonus glaber (Peach, 1883) Pocock, 1911 [Eoscorpius glaber Peach, 1883]
      |  |  |-- †Pseudoarchaeoctonus Kjellesvig-Waering, 1986
      |  |  `-- †Loboarchaeoctonus Kjellesvig-Waering, 1986
      |  `--o †Proscorpiidae Scudder, 1885
      |     |-- †Proscorpius osborni (Whitfield, 1885) Whitfield, 1885 [Palaeophonus osborni Whitfield, 1885]
      |     |-- †Proscorpius Whitfield, 1885
      |     |-- †Archaeophonus Kjellesvig-Waering, 1966
      |     |-- †Waeringoscorpio Størmer, 1970
      |     |-- †Labriscorpio Leary, 1980
      |     `-- †Stoermeroscorpio Kjellesvig-Waering, 1986
      `--o Scorpiones Leach, 1815 [Scorpionoidea Leach, 1815 Euscorpionina Petrunkevitch, 1949; Euscorpiones Petrunkevitch, 1949] (nykyskorpionit)
         |--o †Mesoscorpionina Stockwell, 1996
         |  |-- †Mazonia woodiana Meek & Worthen, 1868 [Mazoniidae Petrunkevitch, 1913; Mazonioidea Petrunkevitch, 1913]
         |  |-- †Mesophonus Wills, 1910
         |  |-- †Heloscorpio Kjellesvig-Waering, 1986
         |  |-- †Phoxiscorpio Kjellesvig-Waering, 1986
         |  |-- †Isobuthus Fric, 1904 [Isobuthidae Petrunkevitch, 1913; Isobuthoidea Petrunkevitch, 1913]
         |  |-- †Eobuthus Fric, 1904
         |  |-- †Eoscorpius carbonarius Meek and Worthen, 1868 [Eoscorpiidae Scudder, 1884]
         |  |-- †Pareobuthus Wills, 1959
         |  |-- †Kronoscorpio Kjellesvig-Waering, 1986
         |  |-- †Paraisobuthus Kjellesvig-Waering, 1986
         |  `-- †Telmatoscorpio Kjellesvig-Waering, 1986
         `--o Neoscorpionina Stockwell, 1996
            |--o †Paleosterni Stockwell, 1996
            |  |-- †Eoctonus miniatus Petrunkevitch, 1913, emend. Petrunkevitch, 1953
            |  |-- †Buthiscorpius buthiformis (Pocock, 1913) Petrunkevitch, 1953 [Anthracoscorpio buthiformis Pocock, 1913]
            |  |-- †Allobuthiscorpius Kjellesvig-Waering, 1986
            |  |-- †Apiscorpio Kjellesvig-Waering, 1986
            |  |-- †Anthracoscorpio juvenis Kušta, 1885 [Cyclophthalmus juvenis (Kušta, 1888)]
            |  |-- †Lichnoscorpius Petrunkevitch, 1949
            |  |-- †Allobuthus Kjellesvig-Waering, 1986
            |  `-- †Coseleyscorpio Kjellesvig-Waering, 1986
            `--o Orthosterni Pocock, 1911
               |?- †Palaeopisthacanthus schucherti Petrunkevitch, 1913
               |?- †Compsoscorpius elegans Petrunkevitch, 1949
               |--o Buthoidea Koch, C.L., 1836 (kenttäskorpioneidensukuiset skorpionit)
               |  |-- Chaerilus [Chaerilidae]
               |  `-- Buthidae Koch, C.L., 1836 (kenttäskorpionit)
               `--+-- Chactoidea
                  `--+-- Scorpionoidea Leach, 1815 (aitoskorpionit)
                     `-- Vaejovoidea

Incertae sedis:

Scorpionida:

        †Brontoscorpio Kjellesvig-Waering, 1972
        †Feistmantelia Fric, 1904
        †Bromsgroviscorpio Kjellesvig-Waering, 1986
        †Titanoscorpio Kjellesvig-Waering, 1986
        †Wattisonia Wills, 1960
        †Palaeomachus Pocock, 1911
        †Tiphoscorpio Kjellesvig-Waering, 1986
        †Eskiscorpio Kjellesvig-Waering, 1986
        †Microlabis sternbergii Corda, 1839 [a member of Isobuthidae Petrunkevitch, 1913 sensu Størmer et al., 1955; Isobuthoidea Petrunkevitch, 1913] Carb. CEu.
        †Hydroscorpius Kjellesvig-Waering, 1986


Scorpiones Hemprich & Ehrenberg, 1837:

        †Alloscorpius granulosus Petrunkevitch, 1949
        †Triconoscorpio americanus Petrunkevitch, 1913
        †Lichnophthalmus pulcher Petrunkevitch, 1949
        †Typhlopisthacanthus mazonensis (Petrunkevitch, 1913) Petrunkevitch, 1949 [Palaeopisthacanthus mazonensis Petrunkevitch, 1913]
        †Typhloscorpius distinctus Petrunkevitch, 1949
        †Europhthalmus longimanus Petrunkevitch, 1949
        †Praearcturus Woodward, 1871
        †Branchioscorpio Kjellesvig-Waering, 1986
        †Acanthoscorpio Kjellesvig-Waering, 1986
        †Gigantoscorpio Størmer, 1963
        †Centromachus Thorell & Lindström, 1885
        †Cyclophthalmus senior Corda, 1835 [Cyclophthalmidae Petrunkevitch, 1913; Cyclophthalmoidea Petrunkevitch, 1913] Carb. Eu.
        †Anthracochaerilus Kjellesvig-Waering, 1986
        †Trachyscorpio Kjellesvig-Waering, 1986
        †Scoloscorpio Kjellesvig-Waering, 1986
        †Garnettius hungerfordi (Elias, 1936) Petrunkevitch, 1953 [Mazonia hungerfordi Elias, 1936]
        †Palaeobuthus Petrunkevitch, 1913
        †Boreoscorpio Kjellesvig-Waering, 1986
        †Leioscorpio Kjellesvig-Waering, 1986
        †Opsieobuthus Kjellesvig-Waering, 1986
        †Pseudobuthiscorpius Kjellesvig-Waering, 1986
        †Petaloscorpio Kjellesvig-Waering, 1986
        †Waterstonia Kjellesvig-Waering, 1986
        †Stenoscorpio Kjellesvig-Waering, 1986
        †Willsiscorpio Kjellesvig-Waering, 1986
        †Spongiophonus Wills, 1947
        †Liassiscorpionides Bode, 1951
        †Mioscorpio Kjellesvig-Waering, 1986

Note:

I'm sorry about the mixed appearance of this page. This is caused by my sources, which are so separate both in time and taxonomy that combining them was very difficult.

Reference(s):

Stockwell, S. A., 1996: Buthoidea.
–iNet: The Tree of Life Project: http://wrbu.si.edu/www/stockwell/phylogeny/buthoidea/buthoidea.html

Stockwell, S. A., 1996: Buthidae.
–iNet: The Tree of Life Project: http://wrbu.si.edu/www/stockwell/phylogeny/buthoidea/buthidae/buthidae.html

Stockwell, S. A., 1996: Chaerilidae.
–iNet: The Tree of Life Project: http://wrbu.si.edu/www/stockwell/phylogeny/buthoidea/chaerilidae/chaerilidae.html

Stockwell, S. A., 1996: Palaeoscorpiones.
–iNet: The Tree of Life Project: http://wrbu.si.edu/www/stockwell/phylogeny/palaeoscorpiones.html

Stockwell, S. A., 1996: Protoscorpiones.
–iNet: The Tree of Life Project: http://wrbu.si.edu/www/stockwell/phylogeny/protoscorpiones.html

Stockwell, S. A., 1996: Scorpionida.
–iNet: The Tree of Life Project: http://wrbu.si.edu/www/stockwell/phylogeny/scorpionida.html

Størmer, L., Petrunkevitch, A. & Hedgpeth, J. W., 1955: Arthropoda 2 - Chelicerata with sections on Pycnogonida and Palaeoisopus. xvii-181
in Moore, R. C. (ed.): Treatise on Invertebrate Paleontology - Part P
–Geological Society of America and University of Kansas Press, Lawrence, 1955