After McKenna & Bell, 1997, de Muizon, Cifelli & Céspedes Paz, 1997 and de Muizon, 1998
<==o †Sparassodonta Ameghino, 1884 [Synonyms] (pussihyeenat; marsupial hyaenas) |?- †Anatherium Ameghino, 1887 [Synonym of †Cladosictis Ameghino, 1887? (McKenna & bell, 1997)] |?- †Plesiofelis Roth, 1903 [Synonym of †Pharsophorus Ameghino, 1897? (McKenna & bell, 1997)] |?- †Pseudonotictis |-- †Sallacyon Villarroel & Marshall, 1982 [Andinogale Hoffstetter & Petter, G., 1983] |--o †Mayulestidae de Muizon, 1994 | |-- †Mayulestes ferox de Muizon, 1994 | `-- †Allqokirus australis Marshall & de Muizon, 1988 `--+-- †Hathliacynidae Ameghino, 1894 [Borhyaenidae: Hathliacyninae Ameghino, 1894 sensu Kirsch, 1977] `--+-- †Borhyaenidae Ameghino, 1894 [incl. Sparassodontidae Roger, 1896; +Conodonictidae Ameghino, F., 1935] (pussihyeenat) `--+--o †Proborhyaenidae Ameghino, 1897 [incl. Arminiheringiidae Ameghino, 1902] | |-- †Arminiheringia acueta Ameghino, 1902 [Dilestes Ameghino, 1902] | |-- †Paraborhyaena Hoffstetter & Petter, G., 1983 | `-- †Proborhyaena gigantea Ameghino, 1897 `--+--o †Prothylacinidae Ameghino, 1894 | |-- †Pseudothylacinus Ameghino, 1902 | |--o †Prothylacynus Ameghino, 1891 [Prothylacocyon Winge, 1923; Napodonictis Ameghino, 1894] | | `-- †P. patagonicus | |-- †Lycopsis longirostris Cabrera, 1927 | |-- †Stylocynus paranensis Mercerat, 1917 | `-- †Pseudolycopsis Marshall, 1976 `--o †Thylacosmilidae Riggs, 1933 sensu Marshall, 1976 (“tikarihammaspussikissat”) |--o †Achlysictis Ameghino, 1891 | |-- †A. acutidens | |-- †A. lelongi Ameghino, 1891 | |-- †A. propampina | `-- †A. purgens Ameghino, 1904 [=? A. pungens] |?- †Amphiproviverra manzaniana |-- †Hyaenodontops chapalmalensis Ameghino, 1908 [Thylacosmilus Riggs, 1933] |-- †Notosmilus pattersoni Kraglievich, J. L., 1960 [Thylacosmilus Riggs, 1933] `--o †Thylacosmilus Riggs, 1933 [incl. ?Notosmilus Kraglievich, 1960; ?Hyaenodontops Ameghino, 1908; Achlysictis Ameghino, 1891; Notosmilus Kraglievich, J. L., 1960; Hyaenodontops Ameghino, 1908] |-- †T. atrox Riggs, 1933 [incl. T. lentis Riggs, 1933??] `-- †T. lentis Riggs, 1933
Reference(s):
- Argot, C., 2004: Evolution of South American mammalian predators (Borhyaenoidea): anatomical and palaeobiological implications.
–Zoological Journal of the Linnean Society: Vol. 140, #4, pp. 487-521 [doi: 10.1111/j.1096-3642.2004.00110.x] - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - de Muizon, C., 1998: Mayulestes ferox, a borhyaenoid (Metatheria, Mammalia) from the early Palaeocene of Bolivia: Phylogenetic and paleobiological implications.
–Geodiversitas: Vol. 20, #1, pp. 19-142
[from abstract] - de Muizon, C., Cifelli, R. L. & Céspedes Paz, R., 1997: The origin of the dog-like borhyaenoid marsupials of South America.
–Nature: Vol. 389, 2 October, pp. 486-489 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631
Special thanks to:
Jos H. M. Dols, for gathering additional information and missing species.