After Nowak, 1991, McKenna & Bell, 1997, and Norris & Musser, 2001
<==o Phalangeridae Thomas, 1888 [taxonomic history] (pussikiipijät) |?- †Cercaertus Burmeister, 1837 [=? Trichosurus Lesson, 1828 (McKenna & Bell, 1997)] |?- †Palaeoptaurus [=? †Palaeopetaurus, jr. synon. of Gymnobelideus (Petauridae)] |-- Ailurops ursinus Wagler, 1830 [non Ailurops Michaelles, 1830] [Ailuropinae Flannery, Archer & Maynes, in Archer, 1987] (karhukuskus) `--o Phalangerinae Thomas, 1888 |--o Strigocuscus Gray, 1862 | |?- †S. reidi Flannery & Archer, 1987 | |?- †S. notialis Flannery, Turnbull, Rich & Lundelius, 1987 | |--o S. celebensis (sulawesinkuskus) | | |?- S. c. feileri Groves, 1987 | | `-- S. c. celebensis | |-- S. pelengensis (“pelengsaartenkuskus”) | |-- S. rothschildi (“obibesarsaartenkuskus”) | |-- S. ornatus (Gray, 1860) (halmahera cuscus) | `-- S. gymnotis (maakuskus) `--+--o Trichosurus Lesson, 1828 [Cercaertus Burmeister, 1837; Psilogrammurus Gloger, 1841; Trichurus Wagner, 1843 (lapsus calami)] (“kettukusut”) | |-- †T. dicksoni Flannery & Archer, 1987 | |-- †T. hamiltonensis Flannery, Turnbull, Rich & Lundelius, 1987 | |--o T. vulpecula (Förster, 1913) (kettukusu) | | |-- T. v. vulpecula (Förster, 1913) | | |-- T. v. raui (Kerr, 1792) | | `-- T. v. fuliginosus | |-- T. arnhemensis [T. vulpecula arnhemensis] (pohjankusu) | `-- T. caninus (Ogilby, 1836) (koirakusu, bobuck) `--+-- Wyulda squamicaudata Alexander, 1919 (suomuhäntäkusu) `--+--o Spilocuscus Gray, 1862 | |-- S. rufoniger (mustatäpläkuskus) | `-- S. maculatus (Desmarest, 1818) [Phalanger maculatus (Desmarest, 1818)] (täpläkuskus) `--o Phalanger Storr, 1780 [Phalangista G. Cuvier & É. Geoffroy Saint-Hilaire, 1795; Coescoes Lacépède, 1799; Balantia Illiger, 1811; Sipalus Fischer de Waldheim, 1813; Cuscus Lesson, 1826; Ceonyx Temminck, 1827] [incl. Eucuscus Gray, 1862] |?- †P. alexandrae Flannery, 1996 |?-o †P. ornatus | |-- †P. o. ornatus | `-- †P. o. matabiru Flannery, 1996 |--o P. orientalis -species-group | |--o P. orientalis (Pallas, 1766) (samoanharmaakuskus; lowland gray cuscus; Samoan Islands cuscus) | | |-- P. o. orientalis (Pallas, 1766) | | |-- P. o. kori Menzies & Pernetta, 1986 | | |-- P. o. timorensis Menzies & Pernetta, 1986 | | `-- P. o. peninsulae Tate, 1945 | |-- P. intercastellanus Thomas, 1895 (pohjanrannikkoharmaakuskus; East Papuan cuscus) | `-- P. mimicus (Thomas, 1922) (etelänrannikkoharmaakuskus; South Papuan cuscus) |--o P. vestitus (silkkikuskus) | |-- P. v. vestitus (silkkikuskus) | `-- P. v. interpositus (steininkuskus) |-- P. carmelitae (vuorikuskus) |-- P. permixtio Menzies & Pernetta, 1986 |--o P. sericeus (silky cuscus) | |-- P. s. sericeus | `-- P. s. occidentalis Menzies & Pernetta, 1986 |-- P. lullulae (Thomas, 1896) (saarikuskus) `-- P. matanim Flannery, 1987 (telefomin cuscus)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Norris, C. A. & Musser, G. G., 2001: Systematic revision within the Phalanger orientalis complex (Diprotodontia, Phalangeridae): a third species of lowland gray cuscus from New Guinea and Australia.
–American Museum Novitates: #3356, pp. 1-20 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629
Special thanks to:
Jos H. M. Dols, for gathering additional information and missing species.