After McKenna & Bell, 1997, with recent species from Nowak, 1991

<==o Protoxerini Moore, 1959
   |--o †Kubwaxerus Cifelli, Ibui, Jacobs & Thorington, 1986; U.Mioc. Af.
   |--o Heliosciurus Trouessart, 1880; U.Pleist.-R. Af. (aurinko-oravat)
   |  |-- H. ruwenzorii (ruwenzorinaurinko-orava)
   |  |-- H. gambianus (gambianaurinko-orava)
   |  |-- H. rufobrachium (redlegged Sun Squirrel)
   |  |-- H. mutabilis ()?
   |  |-- H. undulatus ()?
   |  `-- ?H. punctatus (temminck's Spotted Squirrel)
   |--o Protoxerus Forsyth Major, 1893 [incl. Myrsilus Thomas, 1909 (non Myrsilus Stal, 1866, an insect); Allosciurus Conisbee, 1953] (Oil-palm squirrels, African giant squirrels)
   |  |-- P. (Allosciurus) aubinnii (slendertailed Giant Squirrel)
   |  `-- P. (Protoxerus) stangeri (giant Forest Squirrel)
   `--o Epixerus ebii Thomas, 1909 (African palm squirrels; ebien Squirrel or Temminck's Giant Squirrel)
      |-- E. e. ebii ()?
      |-- E. e. jonesi ()?
      `-- E. e. wilsoni ()?

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