After Repenning, 1983, Winkler & Grady, 1990, Martin, 1987, 1989, 1995, Mou, 1997, McKenna & Bell, 1997, Chaline, et al., 1999, and Martin, Goodwin & Farlow, 2002, with recent species from Nowak, 1991
<==o Arvicolini Gray, 1821 [taxonomic history] (voles; myyrät) |?-+--o †Nebraskomys Hibbard, 1957; L.-U.Plioc. NA. [Arvicolinae incertae sedis sensu McKenna & Bell, 1997] | | |-- †N. mcgrewi Hibbard, 1970 | | `-- †N. rexroadensis Hibbard, 1970 | `--o †Atopomys Patton, 1965; M.Pleist. NA. [Arvicolinae incertae sedis sensu McKenna & Bell, 1997] | |-- †A. texensis Patton, 1965 | `-- †A. salvelinus Zakrzewski, 1975 |--+--o †Promimomys Kretzoi, 1955 [jr. synon. of †Mimomys sensu McKenna & Bell, 1997] | | `-- †P. insuliferus Kowalski, 1956 [Prosomys insuliferus (Kowalski, 1956) sensu Agadjanian & Kowalski, 1978] | `--o †Mimomys Forsyth Major, 1902 [taxonomic history] | |?- †M. orientalis Young, 1935 [taxonomic history] | |?- †M. banchiaonicus Zheng, 1976 | |?- †M. heshuinicus Zheng, 1976 | |?- †M. youhenicus Xue, 1981 | |?- †M. peii Zheng & Li, 1981 | |?- †M. kretzoii | |?- †M. polonicus Kowalski, 1960 | |?- †M. dakotaensis Martin R. A., 1989 | |?- †M. monahani Martin, L., 1972 | |?- †M. irtyshensis Zazhigin, 1980 | |?- †M. reidi | |?- †M. burgondiae | |?- †M. pitymyoides | |?-+-- †M. minor | | `-- †M. medasensis | |?-o †(Borsodia) Janossy & van der Meulen, 1975 | | |-- †B. petenyi | | `-- †B. hungarica | |?-o †(Cseria) Kretzoi, 1959 | | `-- †M. (C.) gracilis | |?-+-- †M. vandermeuleni [M. vandermauleni (lapsus calami)] | | `--+-- †M. occitanus | | |--o †(Mimomys) Forsyth Major, 1902 | | | |-- †M. (M.) davakosi | | | |?- †M. (M.) pliocaenicus | | | `-- †M. (M.) ostramosensis | | `--o †(Kislangia) Kretzoi, 1954 [taxonomic history] | | |-- †M. (K.) adroveri | | `--+-- †M. (K.) cappettai | | `--+-- †M. (K.) gusii | | |-- †M. (K.) ischus | | `-- †M. (K.) rexi | |--o †(Ogmodontomys) Hibbard, 1941; L.-U.Plioc. NA. [treated as a genus by McKenna & Bell, 1997] | | |-- †M. (O.) pipecreekensis Martin, Goodwin & Farlow, 2002 | | |-- †M. (O.) sawrockensis ==> Cosomys primus sensu Chaline, et al., 1999 | | `--o †M. (O.) poaphagus (Hibbard, 1941) | | |-- †M. (O.) poaphagus transitionalis | | `-- †M. (O.) poaphagus poaphagus | `--+-- †M. (?Mimomys) panacensis Mou, 1997 | `--o †(Ophiomys) Hibbard & Zakrzewski, 1967; L.-U.Plioc. NA. [treated as a genus by McKenna & Bell, 1997] | |?- †M. (O.) mcknighti Gustafson | |-- †M. (O.) taylori (Hibbard) | `--+--+-- †M. (O.) magilli | | `--o †M. (Cosomys) R.W. Wilson, 1932; Plioc. NA. [treated as a genus by McKenna & Bell, 1997] | | `-- †M. (C.) primus | `--+-- †M. (O.) meadensis (Hibbard) | `--+-- †M. (O.) parvus (Wilson) | `-- †M. (O.) fricki |--o †Prosomys Shotwell, 1956 [incl. Polonomys Kretzoi, 1959] L.-U.Plioc. Eu., L.Plioc. NA., U.Plioc. As. | |?- †P. insuliferus (Kowalski, 1956) sensu Agadjanian & Kowalski, 1978 [Promimomys insuliferus Kowalski, 1956] | `-- †P. mimus |--o †Kilarcola Kotlia, 1985; U.Plioc.-L.Pleist. As. [Kilarcola Kotlia, 1984 (nomen nudum)] | |-- †K. indicus | `-- †K. kashmiriensis |--o †Hibbardomys Zakrzewski, 1984; U.Plioc. NA. | |-- †H. voorhiesi | |-- †H. fayae | |-- †H. marthae | `-- †H. zakrzewskii Martin, R. A., 1989 |--o †Cromeromys Zazhigin, 1980; U.Plioc. As., L.Pleist. Eu. [Mimomys (Cromeromys) sensu Chaline et al., 1999] | |-- †C. tornensis (Janossy & van der Maulen, 1975) | `-- †C. virginianus (Repenning & Grady, 1988) |--+-- †Mimomys coelodus | `--+-- †Mimomys savini | `--o Arvicola Lacépède, 1799; L.Pleist.-R. As. Eu. (water voles; vesimyyrät) [taxonomic history] | |-- A. sapidus (lännenvesimyyrä) | `-- A. terrestris (vesimyyrä) |-- †Tyrrhenicola Forsyth Major, 1905; |--+-- †Mimomys pusillus | `--+-- †Mimomys newtoni | `--+-- †Mimomys lagurodontoides | `--o “The Allophaiomys -complex” [polyphyletic basal microtines] | |?-+--o Blanfordimys Argyropulo, 1933 [Microtus (Blanfordimys) Argyropulo, 1933 sensu Chaline et al., 1999]; R. As. | | | |-- B. afghanus | | | `-- B. bucharicus | | `-- Microtus (Terricola) Fatio, 1867 (tunnelimyyrät) | `--+?-o Chionomys Miller, 1908 [Microtus (Chionomys) Miller, 1908 sensu Chaline et al., 1999]; M.Pleist.-R. Eu., R. As. | | |-- C. nivalis (lumimyyrä) | | `--+-- C. gud | | `-- C. roberti | |--o Lasiopodomys Lataste, 1887 [tereated as a jr. synon. of Microtus by McKenna & Bell, 1997] | | |-- †L. praebrandti (Erbajeva, 1976) | | |-- †L. probrandti Zheng & Cai, 1991 [cf. Microtus brandtioides Young; Teilhard, 1936; Microtus brandtioides Young; Zheng, 1981; Microtus brandtioides Young; Zheng et al., 1985] | | |-- †L. deceitensis (Guthrie & Matthew, 1971) | | |-- L. brandtii [L. brandtioides (Young)] (brandtinmyyrä) | | |-- L. fuscus | | `-- L. mandarinus | |-- Proedromys bedfordi Thomas, 1911; ?Pleist.-R. EAs. [2 known specimens!] | `-- Microdus Schrank, 1798; U.Plioc.-R. As. Eu. NA., R. Af. CA. (voles; myyrät) [taxonomic history] |--o Volemys Zagorodnyuk, 1990; R. As. | |-- V. clarkei | |-- V. kikuchii | |-- V. millicens | `-- V. musseri |--o †Villanyia Kretzoi, 1956 [incl. †Kulundomys Zazhigin, 1980] U.Plioc. As., U.Plioc.-L.Pleist. Eu. | |-- †V. exilis | |-- †V. laguriformes Erbajeva, 1973 | `-- †V. chinensis [Mimomys? chinensis; Borsonia chinensis sensu Zheng & Cai, 1991] |--+--o Lagurus Gloger, 1841 [Eremiomys Polyakov, 1881; Eremomys Heude, 1898] [incl. †Prolagurus Kormos, 1938; †Lagurodon Kretzoi, 1956; †Laguropsis Kretzoi, 1956 (non Laguropsis Bunge, a plant)] U.Plioc.-R. Eu., L.Pleist.-R. As. | | |-- †L. (Lagurodon) arankae | | `--+--o †(Prolagurus) | | | |-- †L. (P.) leuteus | | | |-- †L. (P.) pannonicus | | | `-- †L. (P.) ternopolitanus | | `--+-- †L. transiens | | `-- L. (Lagurus) lagurus (arosopuli) | `--+--o Eolagurus Argyropulo, 1946; U.Pleist.-R. As., M.Pleist. Eu. | | |-- †E. argyropuloi | | `--+-- E. luteus (yellow steppe lemming; kelta-arosopuli) | | `-- E. przewalskii | `--o Lemmiscus Thomas, 1912 [Lagurus (Lemmiscus) Thomas, 1912; Lemmiscus Davis, 1939] M.Pleist.-R. NA. | `-- L. curtatus (sagebrush vole; maruna-arosopuli) |-- †Jordanomys Haas, 1966; L.Pleist. SWAs. |-- †Kalymnomys W. von Koenigswald, Fejfar & Tchernov, 1992; L.-M.Pleist. SWAs. |--o †Nemausia Chaline & Laborier, 1981; U.Pleist. Eu. | `-- †N. salpetrierensis `-- †Huananomys Zheng, 1992 [†Hexianomys Zheng & Li, 1990 (nomen nudum)] L.-M.Pleist. EAS.
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Chaline, J., Brunet-Lecomte, P., Montuire, S., Viriot, L. & Courant, F., 1999: Anatomy of the arvicoline radiation (Rodentia): palaeogeographical, palaeoecological history and evolutionary data.
–Annales Zoologici Fennici: Vol. 36, pp. 239-267 - Martin, R. A., 1987: Notes on the classification and evolution of some North American fossil Microdus (Mammalia: Rodentia).
–Journal of Vertebrate Paleontology: Vol. 7, #3, pp. 270-283 - Martin, R. A., 1989: Arvicolid rodents of the Early Pleistocene Java local fauna from north-central South Dakota.
–Journal of Vertebrate Paleontology: Vol. 9, #4, pp. 438-450 - Martin, R. A., 1995: A new Middle Pleistocene species of Microdus (Pedomys) from the southern United States, with comments on the taxonomy and early evolution of Pedomys and Pitymys of North America.
–Journal of Vertebrate Paleontology: Vol. 15, #1, pp. 171-186 - Martin, R. A., Goodwin, H. T. & Farlow, J. O., 2002: Late Neogene (Late Hemphillian) rodents from the Pipe Creek sinkhole, Grant County, Indiana.
–Journal of Vertebrate Paleontology: Vol. 22, #1, pp. 137-151 - Martin, R. A., Honey, J. G., Peláez-Campomanes, P., Goodwin, H. T., Baskin, J. A. & Zakrzewski, R. J., 2002: Blancan lagomorphs and rodents of the Deer Park assemblages, Meade County, Kansas.
–Journal of Paleontology: Vol. 76, #6, pp. 1072-1090 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Mou, Y., 1997: A new arvicoline species (Rodentia: Cricetidae) from the Pliocene Panaca Formation, Southeast Nevada.
–Journal of Vertebrate Paleontology: Vol. 17, #2, pp. 376-383 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629 - Repenning, C. A. 1983: Pitymys meadensis Hibbard from the Valley of Mexico and the classification of North American species of Pitymys (Rodentia: Cricetidae)
–Journal of Vertebrate Paleontology: vol. 2, #4,, pp. 471-482. - Winkler, A. J. & Grady, F. V., 1990: The Middle Pleistocene rodent Atopomys (Cricetidae: Arvicolinae) from the Eastern and South-Central United States.
–Journal of Vertebrate Paleontology: Vol. 10, #4, pp. 484-490 - Zheng, S. & Cai, B., 1991: Fossil micromammals from the Donggou section of Dongyaozitou, Yuxian County, Hebei Province.
–Contributions to INQUA XIII: pp. 100-131