After McKenna & Bell, 1997, with Recent species from Nowak, 1991
<==o Castorinae Hemprich, 1820 (beavers; aitomajavat) |?-o †Agnotocastor Stirton, 1935; L.-U.Olig. As. L.Olig. NA. | |-- †A. coloradensis R.W. Wilson, 1949 | |-- †A. galushai Emry, 1972 | |-- †A. montanus (Scott, 1893) [Steneofiber montanus Scott, 1893; Agnotocastor montanus (Scott); Stirton, 1935, and A.E. Wood, 1937] | |-- †A. praetereadens Stirton, 1935 | `-- †A. readingi Korth, 1988 `--+-- †Migmacastor procumbodens Korth & Rybczynski, 2003 `--+--o †Capacikalini L.D. Martin, 1987 | |--o †Capacikala Macdonald, 1963; U.Olig.-L.Mioc. NA., L.Mioc. As. | | |-- †C. gradatus (Cope, 1878) [Steneofiber gradatus Cope, 1878; Palaeocastor gradatus (Cope); Stirton, 1935; Capacikala gradatus (Cope) Macdonald, 1963, Macdonald, 1970, Fremd et al., 1994, and Xu, 1996] | | `-- †C. parvus Xu, 1996 | `?-o †Pseudopalaeocastor L.D. Martin, 1987; L.Mioc. NA. | |-- †P. osmagnus (Xu, 1999) [taxonomic history] | `-- †P. barbouri (Peterson, 1905) [taxonomic history] `--o Castorini Hemprich, 1820 (nykymajavat) |--o †Euhapsina L.D. Martin, 1987 | |--o †Fossorcastor L.D. Martin, 1987; L.Mioc. NA. | | |-- †F. brachyceps (Matthew, 1907) L.D. Martin, 1987 [taxonomic history] | | `-- †F. fossor (Peterson, 1905) [taxonomic history] | `--o †Euhapsis Peterson, 1905; L.Mioc. NA. | |-- †E. breugerorum L.D. Martin, 1987 | |-- †E. ellicottae L.D. Martin, 1987 | |-- †E. luskensis Xu, 1996 | `-- †E. platyceps Peterson, 1905 `--o Castorina Hemprich, 1820 (nykymajavat) |--o †Propalaeocastor Borisoglebskaya, 1967; L.-U.Olig. As. | |-- †P. butselensis | |-- †P. irtyshensis | |-- †P. kazachstanicus | |-- †P. shevyrevae | `-- †P. zaissanensis `--+-- †Palaeomys Kaup, 1832 [Chelodus Kaup, 1832; Chelodon Gloger, 1841 (unjust. emend.); Chaelodus Agassiz, 1842 (lapsus calami)] [incl.? †Chalicomys Kaup, 1832, non Chalicomys Schlosser, 1924 (lapsus calami)] [synonomous status of Palaeomys/Chalicomys is unclear (Korth, 2001)]; U.Olig.-U.Mioc. Eu., M.-U.Olig. As. `--+--o †Palaeocastor Leidy, 1869 [taxonomic history] U.Olig.-U.Mioc. Eu., M.-U.Mioc. As. | |-- †P. nebrascensis (Leidy, 1856) [taxonomic history] | |-- †P. peninsulatus (Cope, 1881) [taxonomic history] | `--o (Capatanka) Macdonald, 1963 | |-- †P. (C.) cankpeopi (MacDonald, 1963) [taxonomic history] | `-- †P. (C.) gaulodon (Matthew, 1907) [taxonomic history] `--+?-o †Hystricops Leidy, 1856 [taxonomic history] ?M.Mioc. As. M.-U. Mioc. As. [treated as Castoroidine by Korth (2017)] | |-- †H. browni Shotwell, 1963 | `-- †H. venustus (Leidy, 1858) [taxonomic history] `--o Castor Linnaeus, 1758 [Fiber Dumeril, 1806 non Fiber G. Cuvier, 1800 (Ondatra)]; Mamcastorus Herrera, 1899] [incl. Sinocastor Young, 1934] U.Mioc.-R. As. Eu. NA. (beavers; majavat) |?- †C. zdanskyi |--o C. fiber Linnaeus, 1758 (Eurasian beaver; majava, euroopanmajava) [8 ssp.] | |-- C. f. fiber (majava; beaver) [Fennoscandia] | |-- C. f. albicus (länsimajava) | |-- C. f. vistulanus | |-- C. f. biellorussieus | |-- C. f. birulai | |-- C. f. pohlei | |-- C. f. tuvinicus | `-- C. f. orientoeuropeas |-- †C. andersoni |-- †C. californicus (kalifornianmajava) `--o C. canadensis Kuhn, 1820 (North American beaver; kanadanmajava) [number and validity of subspecies suspecious] |-- C. c. acadicus Bailey |-- C. c. baileyi Nelson |-- C. c. belugae Taylor (Cook Inlet beaver) |-- C. c. caecator Bangs (Newfoundland beaver) |-- C. c. canadensis Kuhl (Canadian beaver) |-- C. c. concisor |-- C. c. carolinensis Rhoads (Carolina beaver) |-- C. c. duchesnei |-- C. c. frondator Mearns (Sonora beaver) |-- C. c. idoneus |-- C. c. labradorensis |-- C. c. leucodonta Gray (Pacific beaver) |-- C. c. mexicanus Bailey (Rio Grande beaver) |-- C. c. michiganensis Bailey (Woods beaver) |-- C. c. missouriensis Bailey (Missouri River beaver) |-- C. c. pacificus Rhoads (Washington beaver) |-- C. c. pallidus |-- C. c. phaeus Heller (Admiralty beaver) |-- C. c. rostralis |-- C. c. repentinus Goldman (Sonora beaver) |-- C. c. sagittatus |-- C. c. shastensis Taylor (Shasta beaver) |-- C. c. subauratus (California Golden beaver) |-- C. c. taylori Davis |-- C. c. texensis Bailey (Texas beaver)
Reference(s):
- Albright III, L. B., 1996: Insectivorans, rodents, and carnivores of the Toledo Bend Local Fauna: An Arikareean (Earlies Miocene) assemblage from the Texas Coastal Plain.
–Journal of Vertebrate Paleontology: Vol. 16, #3, pp. 458-473 - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Emry, R. J. & Korth, W. W., 1989: Rodents of the Bridgerian (Middle Eocene) Elderberry Canyon local faunaof Eastern Nevada.
–Smithsonian Contributions to Paleobiology: No. 67, pp. iv-14 - Korth, W. W., 2017: A new tribe of castoroidine beavers from the late Arikareean to Hemphillian (Oligocene–Miocene) of western North America.
–Acta Palaeontologica Polonica: Vol. 62, #2, pp. 249-258 [doi: 10.4202/app.00339.2017] - Korth, W. W. & Rybczynski, N., 2003: A new, unusual castorid (Rodentia) from the earliest Miocene of Nebraska.
–Journal of Vertebrate Paleontology: Vol. 23, #3, pp. 667-675 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629