After Nowak, 1991
<==o Colobus Illiger, 1811 [Piliocolobus, Procolobus] (gueretsat) [11] |-- †C. flandrini |-- C. satanas (paholaisgueretsa) |-- C. angolensis (angolangueretsa) |--o C. polykomos (valkohäntägueretsa, mbega) | `-- C. p. polykomos |-- C. vellerosus ()? |-- C. guereza Rüppell, 1935 [incl. †C. polykomos abyssinicus Simons, 1967a] (gueretsa) |--o C. badius [Piliocolobus] (guineanruostegueretsa, gueretsa) | |-- C. b. badius ()? | |-- (†)C. b. waldroni (Waldron's red colobus monkey; waldroninruosteguaretsa) | `-- C. b. temmincki ()? |--o C. pennantii [Piliocolobus] (pennantinruostegueretsa, gueretsa) | |-- C. p. pennantii ()? | |-- C. p. bouvieri () | |-- C. p. preussi (kameruninruostegueretsa, gueretsa)? | |-- C. p. tholloni ()? | `-- C. p. tephrosceles (ugandangueretsa) |--o C. rufomitratus [Piliocolobus] (tananruostegueretsa, gueretsa) | `-- C. r. rufomitratus ()? |-- C. gordonorum [Piliocolobus] (tansaniangueretsa) |-- C. kirkii [Piliocolobus] (sansibaringueretsa) `-- C. verus [Procolobus] (oliiviruostegueretsa (!ruosteguer.))
Reference(s):
- Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629 - Simons, E. L. & Delson, E., 1978: Cercopithecidae and Parapithecidae. 100-119
in Maglio, V. J. & Cooke, H. B. S., (eds.) 1978: Evolution of African Mammals
–Harvard University Press, Cambridge, Massachusetts, London, England, 1978, xiv-641