After McKenna & Bell, 1997, Antoine & Welcomme, 2000 and Antoine, 2002 [Fig. 306]
Alternative phylogeny after Fortelius & Heissig, 1989
<==o †Elasmotheriina Bonaparte, 1845 sensu Antoine, 2002 [Rhinocerotidae: Rhinocerotinae: Elasmotheriini sensu McKenna & Bell, 1997; Rhinocerotidae: Elasmotheriinae: Elasmotheriina Bonaparte, 1845 sensu Prothero & Schoch 1989; Rhinocerotidae: Iranotheriinae] |-- †Kenyatherium bishopi Aguirre & Guérin, 1974 `--+-- †Bugtirhinus praecursor Antoine & Welcomme, 2000 [taxonomic history] |?- †Ougandatherium napakense Guérin & Pickford, 2003 `==o †Caementodon Heissig, 1972 |-- †(Caementodon) oettingenae Heissig, 1972 [taxonomic history] `--+--o †Caementodon (Beliajevina) Heissig, 1974 | |-- †C. (B.) caucasicum (Borissiak, 1935) Heissig, 1974 [taxonomic history] | `-- †C. (B.) fangxianense (Yan, 1979) Antoine, 2002 [taxonomic history] `==o †Hispanotherium Crusafont & Villalta, 1947 |-- †Hispanotherium (Anatolitherium) grimmi (Heissig, 1974) Antoine, 2002 [taxonomic history] `--+-- †Hispanotherium (Hispanotherium) matritense (Lartet, 1864 in Prado, 1864) [taxonomic history] |-- †Hispanotherium (Hispanotherium) corcolense Antoine, Iñigo & Alférez, 2002 [taxonomic history] `--+-- †Hispanotherium (Aegyrcitherium) beonense Antoine, 1997 [taxonomic history] `==o †Procoelodonta Matthew, 1931 |-- †Procoelodonta (Pasalarhinus) tekkayai (Heissig, 1974) Antoine, 2002 [taxonomic history] `--+-- †(Begertherium) borissiaki (Beliajeva, 1971) Beliajeva, 1971 [taxonomic history] `--+-- †(Procoelodonta) mongoliense (Osborn, 1924) Matthew, 1931 [taxonomic history] `--+-- †Huaqingtherium lingtungensis (Zhai, 1978) Huang & Yan, 1983 [taxonomic history] `--+-- †Iranotherium morgani (Mecquenem, 1908) Ringström, 1924 [taxonomic history] `--+-- †Parelasmotherium schansiense Killgus, 1923 [taxonomic history] `--+-- †Sinotherium lagrelii Ringström, 1923 `--o †Elasmotherium Fischer, 1808 [Stereoceros; Enigmatherium Pavlow, 1916] |-- †E. caucasicum Borissiak, 1914 [taxonomic history] `-- †E. sibiricum Fischer, 1808 [taxonomic history]
Reference(s):
- Antoine, P.-O., 2002: Phylogénie et évolution des Elasmotheriina (Mammalia, Rhinocerotidae)
[The Elasmotheriina (Mammalia, Rhinocerotidae): phylogeny and evolution.].
–Publications Scientifiques du Muséum, Mémoires du Muséum national d'Histoire naturelle: Tome 188, 1-359 - Antoine, P.-O. & Welcomme, J.-L., 2000: A new rhinoceros from the Lower Miocene of the Bugti Hills, Baluchistan, Pakistan: the earliest elasmotheriine.
–Palaeontology: Vol. 43, #5, pp. 795-816 - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Guérin, C., 2000: The Neogene Rhinoceroses of Namibia.
–Palaentologica Africana: Vol. 30, pp. 119-138 - Guérin, C. & Pickford, M., 2003: Ougandatherium napakense nov. gen. nov. sp., le plus ancien Rhinocerotidae Iranotheriinae d'Afrique.
[Ougandatherium napakense nov. gen. nov. sp., the earliest Rhinoceratidae Iranotheriinae from Africa].
–Annales de Paléontologie: Vol. 89, #1, pp. 1-35 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629 - Prothero, D. R. & Schoch, R. M. 1989: Classification of the Perissodactyla. 530-537.
in Prothero, D. R. & Schoch, R. M. 1989: The evolution of perissodactyls.
–Oxford University Press., New York. 1989.