After Cifelli & Diaz, 1989 and McKenna & Bell, 1997
<==o †Proterotheriidae Ameghino, 1887 [Proterotheroidea] |--o †Anisolambdinae | |-- †Paranisolambda | |-- †Richardolydekkeria [Lopholambda, Heterolambda] | |-- †Anisolambda [Josepholeidya, Eulambda, Anissolambda] | |-- †Wainka | |-- †Xesmodon | `-- †Protheosodon [Heteroglyphis] `--o †Proterotheriinae |?- †“Proterotherium” dodgei |-- †Deuterotherium |-- †Megadolodus |-- †Eoproterotherium |-- †Prolicaphrium |--o †Prothoatherium Ameghino, 1902 [Licaphrops; Paramacrauchenia; NeodolodusHoffstetter & Soria, 1986] | |-- †P. colombianus (Hoffstetter & Soria, 1986) [Neodolodus colombianus Hoffstetter & Soria, 1986] | `--+-- †P. lacerum | `-- †P. scamnatum |-- †Licaphrium |--o †Diadiaphorus [Coelosoma] | `-- †D. majusculus |--o †Thoatherium | `-- †T. minisculum |-- †Brachytherium [Epitherium, Lophogonodon] |-- †Epecuenia |-- †Diplasiotherium `-- †Proterotherium
Reference(s):
- Cifelli, R. L. & Diaz, J. G., 1989: New remains of Prothoatherium colombianus (Litopterna, Mammalia) from the Miocene of Columbia.
–Journal of Vertebrate Paleontology: Vol. 9, #2, pp. 222-231 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631