After McKenna & Bell, 1997, with Ochotona phylogeny after Ge, Wen, Xia, Zhang, Erbajeva, Huang & Yang, 2013
<==o Ochotonidae Thomas, 1897 (pikas: pikat) |--o †Desmatolagus Matthew & Granger, 1927 [Bohlinotona; Procaprolagus; Desmotolagus (lapsus calami)] | |-- †D. vetustus | `--+-- †D. gobiensis Matthew & Granger, 1927 [Procaprolagus radicidens; Desmotolagus radicidens] | |-- †D. ardynense | |-- †D. robustus | `-- †D. pusillus Teilhard de Chardin, 1926 [Procaprolagus pusillus] |--o †Sinolagomys Bohlin, 1937 [Ochotonolagus] | |?- †S. kansuensis | `-- †S. pachygnathus Li & Qiu, 1980 |-- †Piezodus tomerdingensis Viret, 1929 |-- †Amphilagus ulmensis Tobien, 1974 |?- †Austrolagomys Stromer, in Kaiser, 1926 [a leporid?] |-- †Marcuinomys |--o †Prolagus Pomel, 1853 [Myolagus, Anoema] | |-- †P. crusafonti López Martinez, 1977 | |-- †P. oeningensis (König, 1825) [Anoema oeningensis] | |-- †P. sardus Wagner, 1829 (sardinian pika) [died out during Holocene] | `-- †P. vasconiensis Viret, 1930 |-- †Ptychoprolagus |-- †Heterolagus |--o †Lagopsis Schlosser, 1894 [Opsolagus] | |-- †L. verus (Hensel, 1856) | `-- †L. penai Royo, 1928 |-- †Titanomys von Meyer, 1843 [Platyodon] |--o †Kenyalagomys MacInnes, 1957 | `-- †K. rusingae |--o †Oreolagus Dice, 1917 | |-- †O. colteri | |-- †O. nebrascensis | |-- †O. nevadensis | |-- †O. wallacei | `-- †O. wilsoni |-- †Cuyamalagus dawsoni Hutchison & Lindsay, 1974 |-- †Oklaholmalagus whisenhunti Dalquest et al., 1996 |-- †Albertona |-- †Eurolagus Lopez Martinez, 1977 |--o †Bellatona Dawson, 1961 | `-- †B. yanghuensis Zhou, 1988 |-- †Russellagus vonhofi Storer, 1970 |-- †Gymnesicolagus |--o †Hesperolagomys Clark et al., 1964 | |-- †H. fluviatilis | `-- †H. galbreathi |--o †Alloptox Dawson, 1961 [Metochotona] | |-- †A. anatoliensis | |-- †A. chinghaiensis | `-- †A. gobiensis |-- †Ochotonoides Teilhard de Jardin & Young, 1931 [Prolagomys] |--o Ochotona Link, 1795 [Synonyms] (pikat) | |?- †O. valerotae Erbajeva, Montuire & Chaline, 2001 | |?- †O. polonica | |?- O. lama Michell & Punzo, 1975 | |--o (Conothoa) | | |?- O. gaoligongensis Wang, Gong & Duan, 1988 (gaoligong pika) | | |?- O. nigritia Gong, Wang, Li & Li, 2000 (black pika) | | |--+?- †O. nihewanica Qiu, 1985 [Ochotona lagrelii minor Zheng, 1981] | | | |-- O. erythrotis (Büchner, 1890) (Chinese red-eared pika) | | | `--+-- O. muliensis Pen & Feng, 1962 (muli pika) | | | `-- O. gloveri Thomas, 1922 [Ochotona kamensi] (Glover's pika) | | `--+-- O. forresti Thomas, 1923 (Forrest's pika) | | `--+-- O. rufescens (Gray, 1842) (Afganistan pika, afganistaninpika) | | `--+--+-- O. ladacensis (Günther, 1875) (ladak pika) | | | `--+-- O. koslowi (Buchner, 1894) (Kozlov's pika) | | | `-- O. iliensis Li & Ma, 1986 (ili pika) | | `--+--+-- O. roylei (Ogilby, 1839) [Ochotona angdawai; Ochotona mitchelli; Ochotona nepalensis; Ochotona wardi] (Royle's pika; roylenpika) | | | `-- O. macrotis (Günther, 1875) (Large-eared pika, suurikorvapika) | | `--+-- O. himalayana Feng, 1973 (himalayan pika) | | `-- O. rutila (Severtzov, 1873) (Turkestan red pika; punapika) | `--+--o (Pika) | | |--+-- O. collaris (Nelson, 1893) (collared pika; alaskanpika, kauluspika) | | | `--o O. princeps (Richardson, 1828) (American pika; amerikanpika) | | | |-- O. p. princeps | | | |-- O. p. fenisex | | | |-- O. p. saxatilis | | | |-- O. p. schisticeps | | | `-- O. p. uinta | | `--+--+-- O. argentata Howell, 1928 [Ochotona pallasi helanshanensis Zheng, 1987] (Helan Shan pika, silver pika) | | | `--o O. pallasi (Gray, 1867) (Mongolian Pika; mongolianpika) | | | |-- O. p. pricei | | | |-- O. p. hamica | | | `-- O. p. sundica | | `--+--+-- O. turuchanensis Naumov, 1934 (Turuchan pika) | | | `-- O. alpina (Pallas, 1773) (Alpine pika, Altai Pika; altainpika, altainpiiskujänis) | | `--+-- O. hoffmanni Formozov, Yakhontov & Dmitriev, 1996 [Ochotona alpina hoffmanni Formozov, Yakhontov & Dmitriev, 1996] (Hoffmann's Pika) | | `--o O. hyperborea (Pallas, 1811) (pohjanpika, japaninpika) | | |-- O. h. cinereoflava | | |-- O. h. coreana | | |-- O. h. ferruginea | | |-- O. h. mantchurica | | |-- O. h. normalis | | |-- O. h. uralesis | | |-- O. h. yesoensis | | `-- O. h. yoshikurai | `--o (Ochotona) | |-- O. cansus Lyon, 1907 (gansu pika, gray pika) | `--+-- O. huangensis Matschie, 1908 (Tsing-ling pika) | `--+--o O. pusilla (Pallas, 1769) (Steppe pika; kääpiöpika) | | |-- †O. p. aktogaiensis Savinov, 1981 | | `--+-- †O. p. occidentalis Erbajeva, Montuire & Chaline, 2001 | | |-- †O. p. spelea Owen, 1846 | | |-- O. p. angustifrons Argyropulo, 1932 | | `-- O. p. pusilla (Pallas, 1768) | `--+-- O. thomasi Argyropulo, 1948 (Thomas' pika) | `--+-- O. dauurica (Pallas, 1776) Link, 1795 [Lepus dauuricus Pallas, 1776] (daurianpika) | `--+-- O. thibetana (Milne-Edwards, 1871) (moupin or forest pika) | `--+-- O. nubrica Thomas, 1922 (nubra pika) | `-- O. curzoniae (Hodgson, 1858) (plateau pika, black-lipped pika) |-- †Paludotona etruria `-- †Pliolagomys gigas
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Erbajeva, M. A., Montuire, S. & Chaline, J., 2001: New ochotonids (Lagomorpha) from the Pleistocene of France.
–Geodiversitas: Vol. 23, #3, pp. 395-409 - Ge, D., Wen, Z., Xia, L., Zhang, Z., Erbajeva, M., Huang, C. & Yang, Q., 2013: Evolutionary History of Lagomorphs in Response to Global Environmental Change.
–Plos One: Vol. 8, #4, pp. e59668 [doi: 10.1371/journal.pone.0059668] - Ginsburg, L., 2001: Les faunes de mammifères terrestres du Miocène moyen des Faluns du bassin de Savigné-sur-lathan (France).
[The Middle Miocene land mammals fauna of the Savigné-sur-Lathan basin (France).]
–Geodiversitas: Vol. 23, #3, pp. 381-394 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Meng, J., Hu, Y. & Li, C., 2005: Gobiolagus (Lagomorpha, Mammalia) from Eocene Ula Usu, Inner Mongolia, and comments on Eocene lagomorphs of Asia.
–Palaeontologia Electronica: Vol. 8, #1, pp. 7A: 1-23
[http://palaeo-electronica.org/paleo/2005_1/meng7/issue1_05.htm] - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629 - Tchernov, E., Ginsburg, L., Tassy, P. & Goldsmith, N. F., 1987: Miocene mammals of the Negev (Israel).
–Journal of Vertebrate Paleontology: Vol. 7, #3, pp. 284-310 - Wang, B.-Y. & Qiu, Z.-X., 2004: Discovery of Early Oligocene mammalian fossils from Danghe Area, Gansu, China.
–Vertebrata PalAsiatica: Vol. 42, #2, pp. 130-143 - Zheng, S. & Cai, B., 1991: Fossil micromammals from the Donggou section of Dongyaozitou, Yuxian County, Hebei Province.
–Contributions to INQUA XIII: pp. 100-131