After McKenna & Bell, 1997, with Recent species from Nowak, 1991
<==o Cynopterini
|--o Nyctimenina Miller, 1907 ()?
| |-- Paranyctimene raptor ()?
| `--o Nyctimene (putkikuonolepakot)
| |-- N. minutus (pikkuputkikuonolepakko)
| |-- N. albiventer ()?
| |-- N. draconilla ()?
| |-- N. robinsoni ()?
| |-- N. cephalotes ()?
| |-- N. vizcaccia ()?
| |-- N. masalai ()?
| |-- N. rabori ()?
| |-- N. malaitensis ()?
| |-- N. major (isoputkikuonolepakko)
| |-- N. sanctacrucis ()?
| |-- N. cyclotis ()?
| |-- N. aello ()?
| |-- N. celaeno ()?
| `-- ?N. certans ()?
`--o Cynopterina
|--o Cynopterus ()?
| |-- C. brachyotis ()?
| |-- C. minor ()?
| |-- C. archipelagus ()?
| |-- C. horsfieldi ()?
| |-- C. sphinx (lyhytkuonohedelmälepakko)
| |-- C. titthaecheilus ()?
| `-- ?C. nusatenggara ()?
`--+--o Megaerops ()?
| |-- M. niphanae ()?
| |-- M. ecaudatus ()?
| |-- M. kusnotoi ()?
| `-- M. wetmorei ()?
`--+--o Ptenochirus ()?
| |-- P. minor ()?
| `-- P. jagori ()?
|-- Dyacopterus spadiceus ()?
`--+-- Chironax melanocephalus (hattuhedelmälepakko)
`--+--o Thoopterus nigrescens ()?
| |-- T. n. nigrescens ()?
| `-- T. n. latidens ()?
`--+-- Sphaerias blanfordi ()?
`--+-- Balionycteris maculata (täpläpoimuhedelmälepakko)
`--+-- Aethalops alecto (kääpiöhedelmälepakko)
`--+-- Penthetor lucasi ()?
`--+-- Latidens salimalii ()?
`--+-- Alionycteris paucidentata ()?
`--+-- Otopteropus cartilagonodus ()?
`-- Haplonycteris fischeri ()?
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631
- Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii
- Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629