After McKenna & Bell, 1997, with Recent species from Nowak, 1991
<==o Cynopterini |--o Nyctimenina Miller, 1907 ()? | |-- Paranyctimene raptor ()? | `--o Nyctimene (putkikuonolepakot) | |-- N. minutus (pikkuputkikuonolepakko) | |-- N. albiventer ()? | |-- N. draconilla ()? | |-- N. robinsoni ()? | |-- N. cephalotes ()? | |-- N. vizcaccia ()? | |-- N. masalai ()? | |-- N. rabori ()? | |-- N. malaitensis ()? | |-- N. major (isoputkikuonolepakko) | |-- N. sanctacrucis ()? | |-- N. cyclotis ()? | |-- N. aello ()? | |-- N. celaeno ()? | `-- ?N. certans ()? `--o Cynopterina |--o Cynopterus ()? | |-- C. brachyotis ()? | |-- C. minor ()? | |-- C. archipelagus ()? | |-- C. horsfieldi ()? | |-- C. sphinx (lyhytkuonohedelmälepakko) | |-- C. titthaecheilus ()? | `-- ?C. nusatenggara ()? `--+--o Megaerops ()? | |-- M. niphanae ()? | |-- M. ecaudatus ()? | |-- M. kusnotoi ()? | `-- M. wetmorei ()? `--+--o Ptenochirus ()? | |-- P. minor ()? | `-- P. jagori ()? |-- Dyacopterus spadiceus ()? `--+-- Chironax melanocephalus (hattuhedelmälepakko) `--+--o Thoopterus nigrescens ()? | |-- T. n. nigrescens ()? | `-- T. n. latidens ()? `--+-- Sphaerias blanfordi ()? `--+-- Balionycteris maculata (täpläpoimuhedelmälepakko) `--+-- Aethalops alecto (kääpiöhedelmälepakko) `--+-- Penthetor lucasi ()? `--+-- Latidens salimalii ()? `--+-- Alionycteris paucidentata ()? `--+-- Otopteropus cartilagonodus ()? `-- Haplonycteris fischeri ()?
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629
