After Bergmans, 1997 [from Giannini & Simmons, 2003]
<==o Cynopterinae Andersen, 1912 |--o Cynopterus ()? | |-- C. brachyotis ()? | |-- C. minor ()? | |-- C. archipelagus ()? | |-- C. horsfieldi ()? | |-- C. sphinx (lyhytkuonohedelmälepakko) | |-- C. titthaecheilus ()? | `-- ?C. nusatenggara ()? |--o Megaerops ()? | |-- M. niphanae ()? | |-- M. ecaudatus ()? | |-- M. kusnotoi ()? | `-- M. wetmorei ()? |--o Ptenochirus ()? | |-- P. minor ()? | `-- P. jagori ()? |-- Dyacopterus spadiceus ()? |-- Chironax melanocephalus (hattuhedelmälepakko) |--o Thoopterus nigrescens ()? | |-- T. n. nigrescens ()? | `-- T. n. latidens ()? |-- Sphaerias blanfordi ()? |-- Balionycteris maculata (täpläpoimuhedelmälepakko) |-- Aethalops alecto (kääpiöhedelmälepakko) |-- Penthetor lucasi ()? |-- Latidens salimalii ()? |-- Alionycteris paucidentata ()? |-- Otopteropus cartilagonodus ()? `-- Haplonycteris fischeri ()?
Reference(s):
- Butler, P. C., 1978: Insectivora and Chiroptera. 56-68
in Maglio, V. J. & Cooke, H. B. S., (eds.) 1978: Evolution of African Mammals, 1.
–Harvard University Press, Cambridge, Massachusetts, London, England, 1978, xiv-641 - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Giannini, N. P. & Simmons, N. B., 2003: A phylogeny of megachiropteran bats (Mammalia: Chiroptera: Pteropodidae) based on direct optimization analysis of one nuclear and four mitochondrial genes.
–Cladistics: Vol. 19, pp. 496-511 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631 - Nowak, R. M., 1991: Walker's Mammals of the World; Part 1.
–The Johns Hopkins University Press, Baltimore and London, 1991, xlviii-642-lxiii - Nowak, R. M., 1991: Walker's mammals of the World; Part 2.
–The Johns Hopkins University Press, Baltimore and London, 1991, xii-643-1629