<==o UNGULATOMORPHA
   |-- †Aspanlestes aptap [paraphyletic Zhelestidae]
   `--+-- †Avitotherium utahensis [paraphyletic Zhelestidae]
      `--+-- †Alostera saskatchewanensis [paraphyletic Zhelestidae]
         `--+--o "Zhelestinae"
            |  |-- †Zhelestes temirkazyk [paraphyletic Zhelestidae]
            |  `--o †Parazhelestes [paraphyletic Zhelestidae]
            |     |-- †P. robustus [paraphyletic Zhelestidae]
            |     `-- †P. sp. [paraphyletic Zhelestidae]
            `--+-- †Unnamed genus et species [paraphyletic Zhelestidae]
               `--o UNGULATA
                  |-- †Ocyclaenidae [Arctocyonidae: Ocyclaeninae]
                  `--+-- ?†Arctocyonidae  [Arctocyonidae: Arctocyoninae]
                     |-- ?†Paroxyclaenidae
                     |-- ?†Tricuspiodontidae
                     |--+-- ?†Mioclaenidae
                     |  `-- ?†Periptychidae
                     |--o TUBULIDENTATA (putkihampaiset)
                     |  `-- Orycteropodidae (Maasiat)
                     |--o †ARCTOSTYLOPIDA
                     |  `-- †Arctostylopidae
                     |--+-- †Triisodontidae [Arctocyonidae: Triisodontinae]
                     |  `--o "ARTIODACTYLOMORPHA" (sorkkaeläintensukuiset ungulaatit)
                     |     |-- CETE (valasmaiset ungulaatit)
                     |     `-- ARTIODACTYLA (sorkkaeläimet)
                     |--o ALTUNGULATA [Paenungulata]
                     |  |--+-- †Loxolophidae [Arctocyonidae: Loxolophinae]
                     |  |  `--+-- †Phenacolophidae
                     |  |     `-- TETHYTHERIA (norsunsukuiset ungulaatit)
                     |  `--+-- †Meniscotheriidae
                     |     `--+-- †Phenacodontidae
                     |        `-- "PERISSODACTYLOMORPHA" (kavioeläintensukuiset ungulaatit)
                     `--+-- †Didolodontidae
                        `-- MERIDIUNGULATA (etelä-amerikkalaiset ungulaatit)