After Lihoreau & Ducrocq, 2007, with genera et species from Black, 1978, McKenna & Bell, 1997, Ducrocq et al., 2000, and Hooker & Thomas, 2001
<==o †Anthracotheriidae Leidy, 1869 non Gill, 1872 [incl. Bothriodontinae; Anthracotheriinae; ?Hippopotamidae (Lihoreau & Ducrocq, 2007)] |--o †Siamotherium Suteenthorn et al., 1988 | |-- †S. krabiense Suteenthorn et al., 1988 | `-- †S. pondaungensis Ducrocq et al., 2000 [=? Pakkokuhyus lahirii Tsubamoto et al., 2002] |-- †Anthracohyus choeroides Pilgrim & Cotter, 1916 `--+--o †Anthracotheriinae Leidy, 1869 sensu Lihoreau & Ducrocq, 2007 | |--o †Heptacodon Marsh, 1894 [Anthracotherium (in partim); Octacodon Marsh, 1894] | | |-- †H. curtus Marsh, 1894 | | |-- †H. occidentale Osborn & Wortman, 1894 | | |-- †H. pellionis Storer, 1983 | | `-- †H. yeguaensis Holroyd, 2002 | `--+-- †Prominatherium dalmatinum (von Meyer, 1854) Teller, 1884 | `--o †Anthracotherium Cuvier, 1822 [Bugtitherium Pilgrim, 1908; Anthracothema Pilgrim, 1928; ?Heothema Tang, 1978; ?Huananothema Tang, 1978] | |-- †A. alsaticum Cuvier, 1822 | |-- †A. bimonsvialensemagnum Colpe-Posse, 1971 | |-- †A. bugtiense Pilgrim, 1907 | |-- †A. bumbachense Stehlin, 1910 | |-- †A. chaimanei Ducrocq, 1999 | |-- †A. cuvieri Gadry, 1873 | |-- †P. crassum Pilgrim & Cotter, 1916 | |-- †A. hippoideum Rütimeyer, 1857 | |-- †A. illyricum Teller, 1886 | |-- †A. kwablianicum gabunia, 1964 | |-- †A. seckbachense Kinkelin, 1884 | |-- †A. pangan Pilgrim & Cotter, 1916 | |-- †A. valdense Kowalevski, 1876 | `--+-- †A. monsvialense De Zigno, 1888 | `-- †A. magnum Cuvier, 1822 [type] `--+--o †Microbunodontinae Lihoreau & Ducrocq, 2007 | `==o †Anthracokeryx Pilgrim & Cotter, 1916 [Anthracotherium (in partim); Anthracohyus (in partim); Anthracosenex Zdansky, 1930; ?†Probrachyodus Xu, 1962] [paraphyletic sensu Lihoreau & Ducrocq (2007)] | |-- †A. birmanicus Pilgrim & Cotter, 1916 [type] | |-- †A. gungkangensis Qiu, 1977 [A. kwangsiensis Qiu, 1977] | |-- †A. sinensis Zdansky, 1930 | |-- †A. tenuis Pilgrim & Cotter, 1916 | |-- †A. ulnifer Pilgrim, 1928 | `--+-- †A. thailandicus Ducrocq, 1999 | `--o †Microbunodon Depéret, 1908 [Anthracotherium (in partim); Sus (in partim); Microselenodon Depéret, 1908; Rhagatherium (in partim)] | |-- †M. minimum (Cuvier, 1822) | |-- †M. silistrensis (Pentland, 1828) | `-- †M. milaensis Lihoreau et al., 2004 `--o †“Bothriodontinae” Scott, 1940 sensu Lihoreau & Ducrocq, 2007 |?- †Kukusepasutanka schultzi Macdonald, 1956 |?-o †Telmatodon Pilgrim, 1907 [Ancodus (in partim); Brachyodus (in partim); Gonotelma (in partim)] | |-- †T. bugtiensis Pilgrim, 1907 | `-- †T. orientale Forster-Cooper, 1924 |?- †Ulausuodon parvus Hu, 1963 |?- †Parabrachyodus hyopotamoides (Lydekker, 1883) Forster-Cooper, 1915 [Anthracotherium (in partim); Hyopotamus (in partim); Brachyodus (in partim); Telmatodon (in partim)] `--+--+-- †Qatraniodon parvus (Andrews, 1906) Ducrocq, 1997 [Ancodon (in partim); Brachyodus (in partim); Bothriogenys (in partim)] | `--o †Bothriogenys Schmidt, 1913 [Ancodon (in partim); Brachyodus (in partim)] | |-- †B. fraasi Schmidt, 1913 [Brachyodus (Bothriogenys) fraasi Schmidt, 1913] | |-- †B. andrewsi Schmidt, 1913 | |-- †B. gorringei (Andrews & Beadnell, 1902) | |-- †B. rugulosus Schmidt, 1913 | `-- †B. orientalis Ducrocq, 1997 `--+--o †Brachyodus Depéret, 1895 [Masritherium Fourtau, 1918; Bothriogenys (in partim); Anthracotherium (in partim); Hypopotamus (in partim); Sihongotherium Liu & Zhang, 1993] | |-- †B. onoideus (Gervais, 1959) [type] | |-- †B. depereti (Fourtau, 1918) [Masritherium depereti Fourtau, 1918] | |-- †B. aequatorialis MacInnes, 1951 [Masritherium aequitorialis (MacInnes, 1951)] | `-- †B. mogharensis Pickford, 1991 `--+--+--o †Aepinacodon Troxell, 1921 [Hyopotamus (in partim); Ancodus (in partim); Ancodon (in partim); Bothriodon (in partim)] | | |-- †A. americanus (Leidy, 1856) | | `-- †A. deflectus (Marsh, 1890) [type] | `--o †Bothriodon Aymard, 1846 [Ancodon (in partim); Ancodus (in partim); Anthracotherium (in partim); Hyopotamus (in partim)] | |-- †B. advena Russell, 1978 | |-- †B. aymardi (Pomel, 1847) | |-- †B. rostratus (Scott, 1894) | |-- †B. leptorynchus Aymard, 1846 | `-- †B. velaunus (von Meyer, 1832) [type] `==o Elomeryx Marsh, 1894 [Hyopotamus (in partim); Xiphodon (in partim); Brachyodus (in partim); Ancodus (in partim); Microbunodon (in partim); Bunobrachyodus (in partim); Heptacodon (in partim); Bothriodon (in partim)] [paraphyletic sensu Lihoreau & Ducrocq, 2007] |-- †E. crispus (Gervais, 1849) [type] |-- †E. armatus Marsh, 1894 |-- †E. cluai (Depéret, 1906) |--o †Bakalovia Nikolov & Heissig, 1985 [Elomeryx (in partim)] | |-- †B. palaeopontica (Nikolov, 1967) | `-- †B. astica (Nikolov, 1967) `--+-- †E. borbonicus (Geais, 1934) |--o †Arretotherium Douglass, 1901 | |-- †A. acridens Douglass, 1901 [type] | |-- †A. fricki Macdonald & Schultz, 1956 | `-- †A. leptodus (Matthew, 1909) `--+--o †Afromeryx Pickford, 1991 [Brachyodus (in partim); Bothriogenys (in partim); Ancodus (in partim); Hyoboops (in partim); Masritherium (in partim); Gelasmodon (in partim)] | |-- †A. zelteni Pickford, 1991 | `-- †A. africanus (Andrews, 1899) [Bothriogenys africanus (Andrews, 1899)] `--+--+-- †Sivameryx palaeindicus (Lydekker, 1877) Lydekker, 1883 [Hyoboops palaeindicus (Lydekker, 1877); †Hyoboops africanus (Andrews, 1914); Gonotelma shabhazi Pilgrim, 1908] [Choeromeryx; Hyopotamus (in partim); Hyoboops (in partim); Merycops; Ancodon (in partim); Hemimeryx (in partim); Brachyodus (in partim); Rhagatherium (in partim); Gonotelma (in partim)] | `-- †Hemimeryx blandfordi Lydekker, 1883 [Ancodus (in partim); Gelasmodon] `--+--o †Libycosaurus Bonarelli, 1947 [Anthracotherium (in partim); Gelasmodon (in partim);Merycopotamus (in partim)] | |-- †L. algeriensis Ducrocq et al., 2001 | |-- †L. anisae (Black, 1972) [Merycopotamus anisae Black, 1972] | `-- †L. petrocchii Bonarelli, 1947 [Merycopotamus petrocchii (Bonarelli, 1947)] `--+--o †Merycopotamus Falconer & Cautley, 1847 [Hippopotamus (in partim); Hemimeryx (in partim)] | |-- †M. nanus Flaconer, 1868 (ex pusillus) [Hemimeryx pusillus] | `--+-- †M. medioximus Lihoreau et al., 2004 | `-- †M. dissimilis Falconer & Cautley, 1847 `?- Hippopotamidae Gray, 1821 (hippos; virtahevot)
Anthracotheriidae incertae sedis [taxa not treated by Lihoreau & Ducrocq, 2007]:
†“Rhagatherium” aegypticum Andrews, 1906
†Hyoboops Trouessart, 1904 [Merycops]
†H. moneyi (Fourtau, 1918)
†H. sp [Djebel Zelten, Libya] [Black, 1978]
?†“Anthracotherium” punjabiensis
†Probrachyodus
†Atopotherium Ducrocq et al., 1996
†Diplopus
†Aepinacodon
†Pachychoerops
†Bunobrachyodus
†Anthracochoerus Dal Piaz, 1930
†A. stehlini
†A. fabianii
†Gonotelma
†Choeromeryx
Reference(s):
- Black, C. C., 1978: Anthracotheriidae. 423-434.
in Maglio, V. J. & Cooke, H. B. S., (eds.) 1978: Evolution of African mammals.
–Harvard University Press, Cambridge, Massachusetts & London, England, 1978, xiv-641 - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Ducrocq, S., Soe, A. N., Aung, A. K., Benammi, M., Bo, B., Chaimanee, Y., Tun, T., Thein, T. & Jaeger, J.-J., 2000: A new anthracotheriid artiodactyl from Myanmar, and the relative ages of the Eocene anthropoid primate-bearing localities of Thailand (Krabi) and Myanmar (Pondaung).
–Journal of Vertebrate Paleontology: Vol. 20, #4, pp. 755-760 - Hooker, J. J. & Thomas, K. M., 2001: A new species of Amphirhagatherium (Choeropotamidae, Artiodactyla, Mammalia) from the Late Eocene Headon Hill Formation of southern England and phylogeny of endemic european "anthracotheroids".
–Palaeontology: Vol. 44, #5, pp. 827-853 - Lihoreau, F. & Ducrocq, S., 2007: Family Anthracotheriidae. 89-105.
in Prothero, D. R. & Foss, S. E., (eds.) 2007: The evolution of artiodactyls.
–The Johns Hopkins University Press, Baltimore, 2007, xii-367 - McKenna, M. C. & Bell, S. K., (eds.) 1997: Classification of mammals – above the species level.
–Columbia University Press, New York, 1997, xii-631