After Bell, 1997, Creisler [& Poling], 2000, Polcyn & Bell, 2005, Cuthbertson, Mallon, Campione & Holmes, 2007, and Konishi & Caldwell, 2011
<==o †Russelosaurina [Tylosaurinae & Plioplatecarpinae] |--o †Tethysaurinae Makádi, Caldwell & Osi, 2012 [“Russellosaurinae”] | |-- †Pannoniasaurus inexpectatus Makádi, Caldwell & Osi, 2012 | `--+-- †Tethysaurus nopcsai Bardet, Pereda-Suberbiola & Jalil, 2003 | `--+-- †Yaguarasaurus columbianus Paramo, 1994 | `-- †Russellosaurus coheni Polcyn & Bell, 2005 `--+--o †Tylosaurinae (Williston, 1897) sensu Bell, 1997, Polcyn & Bell, 2005 | |?- †Lakumasaurus antarcticus Novas et al., 2002 | `--o †Tylosaurus Marsh, 1872 sensu lato [taxonomic history] | |-- †“Tylosaurus” kansasensis Everhart, 2005 [was “Tylosaurus n. sp. under description” (Bell, 1997)] | |-- †“Tylosaurus” ivoensis (Persson, 1963) U. Cret. NEu. [†Mosasaurus ivoensis Persson, 1963] | `--+-- †“Tylosaurus” nepaeolicus (Cope, 1874) | `--+-- †Tylosaurus proriger (Cope, 1869); Marsh, 1872 [taxonomic history] | `--+?- †Taniwhasaurus oweni Hector, 1874 [taxonomic history] | `--o †Hainosaurus Dollo, 1885 | |-- †H. bernardi Dollo, 1885 | `-- †H. pembinensis Nicholls, 1988 `--o †Plioplatecarpinae (Dollo, 1884) sensu Bell, 1997, Polcyn & Bell, 2005 [Plioplatecarpidae Dollo, 1884] |?- †Igdamanosaurus aegyptiacus (Zdansky, 1935) [Globidens aegyptiacus Zdansky, 1935] |--o †Ectenosaurus Russell, 1967 | |-- †E. clidastoides (Merriam, 1894) | `-- †E. sp. `--+-- †Angolosaurus bocagei Telles-Antunes, 1964 [“Platecarpus" bocagei (Telles-Antunes, 1964)] `--+--o †Selmasaurus Wright & Shannon, 1988 | |-- †S. russelli Wright & Shannon, 1988 | `-- †S. johnsoni Polcyn & Everhart, 2008 `--+-- †Plesioplatecarpus planifrons (Cope, 1874) Konishi & Caldwell, 2011 [Clidastes planifrons Cope, 1874; Platecarpus planifrons Williston, 1898; Platecarpus planifrons (Cope); Konishi & Caldwell, 2007] `--+-- †Platecarpus tympaniticus (Cope, 1869) Cope, 1869 [taxonomic history] `==o †Latoplatecarpus Konishi & Caldwell, 2011 |-- †L. willistoni Konishi & Caldwell, 2011 (type) `--+-- †L. nichollsae (Cuthbertson, Mallon, Campione & Holmes, 2007) [Plioplatecarpus nichollsae Cuthbertson, Mallon, Campione & Holmes, 2007] L.Cret. NAm. |-- †“Platecarpus” somenensis Thevenin, 1896 `--o †Plioplatecarpus Dollo, 1882 [Oterognathus Dollo, 1889] |?- †“Mosasaurus” scanicus |?- †P. crassartus |?- †P. depressus [Baptosaurus fraternus Cope, 1869-70] |-- †P. sp. [UNO 861-2] [Holmes, 1996] [Late Cretaceous, Bearpaw Formation, North America] `--+-- †P. primaevus Russell, 1967 `--+-- †P. houzeaui (Dollo, 1889) [Oterognathus houzeaui Dollo, 1889] `-- †P. marshii Dollo, 1882
Reference(s):
- Bell, G. L., 1997: A phylogenetic revision of North American and Adriatic Mosasauroidea. 293-332
in Callaway, J. M. & Nicholls, E. L., (eds.), 1997: Ancient marine reptiles.
–Academic Press, San Diego, 1997, pp. xlvi - 501 - Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Caldwell, M. W., Holmes, R. B., Bell Jr., G. L. & Wiffen, J., 2005: An unusual tylosaurine mosasaur from New Zealand: a new skull of Taniwhasaurus oweni (Lower Haumurian; Upper Cretaceous).
–Journal of Vertebrate Paleontology: Vol. 25, #2, pp. 393-401 - Creisler, B. [& Poling, J.], 2000: Mosasauridae translation and pronunciation guide.
–iNet: Dinosauria.com: http://www.dinosauria.com/dml/names/mosa.html - Cuthbertson, R. S., Mallon, J. C., Campione, N. E. & Holmes, R. B., 2007: A new species of mosasaur (Squamata: Mosasauridae) from the Pierre Shale (lower Campanian) of Manitoba.
–Canadian Journal of Earth Sciencies: Vol. 44, #5, pp. 593-606 [doi: 10.1139/E07-006] - Holmes, R., 1996: Plioplatecarpus primaevus (Mosasauridae) from the Bearpaw Formation (Campanian, Upper Cretaceous) of the North American Western Interior Seaway.
–Journal of Vertebrate Paleontology: Vol. 16, #4, pp. 673-687 - Konishi, T. & Caldwell, M. W., 2007: New specimens of Platecarpus planifrons (Cope, 1874) (Squamata: Mosasauridae) and a revised taxonomy of the genus.
–Journal of Vertebrate Paleontology: Vol. 27, #1, pp. 59-72 [doi: 10.1671/0272-4634(2007)27[59:nsoppc]2.0.co;2] - Konishi, T. & Caldwell, M. W., 2011: Two New Plioplatecarpine (Squamata, Mosasauridae) Genera from the Upper Cretaceous of North America, and a Global Phylogenetic Analysis of Plioplatecarpines.
–Journal of Vertebrate Paleontology: Vol. 31, #4, pp. 754-783 [doi: 10.1080/02724634.2011.579023] - Lindgren, J. & Siverson, M., 2003: A possible global mosasaur extinction event at the lower/upper Campanian boundary. [an abstract]
–GFF: Vol. 125, pp. 107 - Makádi, L., Caldwell, M. W. & Osi, A., 2012: The First Freshwater Mosasauroid (Upper Cretaceous, Hungary) and a New Clade of Basal Mosasauroids.
–Plos One: Vol. 7, #12, pp. e51781 [doi: 10.1371/journal.pone.0051781] - Everhart, M. J., 2005: Earliest record of the genus Tylosaurus (Squamata; Mosasauridae) from the Fort Hays Limestone (Lower Coniacian) of western Kansas.
–Transactions of the Kansas Academy of Science: Vol. 108, #3/4, pp. 149-155 - Everhart, M. J., 2005: Tylosaurus kansasensis, a new species of tylosaurine (Squamata, Mosasauridae) from the Niobrara Chalck of western Kansas, USA.
–Netherlands Journal of Geosciences – Geologie en Mijnbouw: Vol. 84, #3, pp. 231-240 - Polcyn, M. J. & Bell Jr., G. L., 2005: Russellosaurus coheni n. gen., n. sp., a 92 million-year-old mosasaur from Texas (USA), and the definition of the parafamily Russellosaurina. –Netherlands Journal of Geosciences - Geologie en Mijnbouw: Vol. 84, #3 [from abstract]
- Storrs, G. W., Arkhangel'skii, M. S. & Efimov, V. R., 2000: Mezosoic marine reptiles of Russia and other former Soviet republics. 187-210
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000 xxxix-696 - Wright, K. R. & Shannon, S. W., 1988: Selmasaurus russelli, a new plioplatecarpine mosasaur (Squamata, Mosasauridae) from Alabama.
–Journal of Vertebrate Paleontology: Vol. 8, #1, pp. 102-107