Mikko's Phylogeny Archive

Anguidae: Glyptosaurinae – glyptosaurine lizards

After Sullivan, 1986, Zonneveld, Gunnell & Bartels, 2000, Augé, 2003, Sullivan & Augé, 2006, and Augé & Sullivan 2006

<==o †Glyptosaurinae Marsh, 1872 sensu Sullivan, 1986
   |?- †Paraxestops stehlini Hoffstetter, 1962; U.Eoc. Eu.
   |--o †Odaxosaurus [†Pancelosaurus Meszoely 1970] [‘Odaxosaurini’ Sullivan 1979]
   |  |-- †O. jepseni Meszoley et al., 1978
   |  |-- †O. niger Gilmore, 1928
   |  |-- †O. piger Gilmore, 1928 [Odaxosaurus obliquus Gilmore 1928; Pancelosaurus piger; Peltosaurus piger]
   |  `-- †O. priscus Gao & Fox, 1996
   `--o †‘Melanosaurini’ Sullivan 1979 [prob. paraphyletic (Augé & Sullivan, 2006)]
      |-- †Proxestops silberlingii (Gilmore, 1938) Gauthier, 1982 [P. jepseni Gauthier, 1982]
      `--+?- †Xestops vagans (Marsh, 1873) Cope, 1873 sensu Keller et al., 1991 [Oreosaurus] Eoc. NA.
         |?-o †Peltosaurus Cope, 1872
         |  |?- †P. minimus
         |  |-- †P. floridanus Vanzolini, 1952
         |  |-- †P. granulosus Cope, 1873 [Peltosaurus abbotti Gilmore, 1928]
         |  `-- †P. macrodon Brattstrom, 1955
         |--+-- †Arpadosaurus gazinorum Meszoely, 1970; Eoc. CEu.
         |  |-- †Melanosaurus maximus Gilmore, 1928; Eoc. NAm.
         |  |-- †Paraplacosauriops quercyi (Filhol, 1882) Augé & Sullivan, 2006 [†Plestiodon quercyi Filhol, 1882; Diploglossus cadurcensis De Stefano, 1903; Placosaurus leenhardti Leenhardtm 1926; Placosaurus quercyi Hoffstetter, 1944; Placosaurus quercyi (Filhol, 1882) Sullivan & Augé, 2006]
         |  `--o †Placosauriops Kuhn, 1940; Eoc. Eu. [taxonomic history]
         |     |-- †P. weigelti (Kuhn, 1940) [taxonomic history]
         |     `?- ?†Plecosauriops indet. (Augé, 2003)
         `--o †Glyptosaurini Sullivan, 1979
            |?- †Proglyptosaurus huerfanensis [Eoglyptosaurus donohoei Sullivan, 1979 (in partim)] [AMNH 7431]
            |--+--o †Glyptosaurus Marsh, 1871 [taxonomic history]
            |  |  |-- †G. anceps Marsh, 1871
            |  |  `-- †G. sylvestris Marsh, 1871 [taxonomic history]
            |  `?-o †Paraglyptosaurus Sullivan, 1979 [=? Glyptosaurus (Sullivan, 1986)]
            |     |?- †P. yatkolai Sullivan, 1979
            |     |-- †P  princeps (Marsh, 1872) [= Glyptosaurus sylvestris Marsh, 1871 (sensu Sullivan, 1986)]
            |     `-- †P. hillsi Gilmore, 1928
            `--+-- †Helodermoides tuberculatus Douglass, 1903
               `--o †Placosaurus Gervais, 1848-52 [taxonomic history]
                  |?- †P. cayluxi (nomen dubium)
                  |?- †P. galliae (nomen dubium)
                  |?- †P. waltheri (nomen dubium)
                  |?- †P. margariticeps (Gervais, 1876) (nomen dubium, type lost) [taxonomic history]
                  |?- †P.? europaeus (Fihol, 1876) [taxonomic history]
                  |-- †P. rugosus Gervais, 1848-52
                  |-- †P. mongoliensis (Sullivan, 1979) [taxonomic history]
                  `-- †P. estesi Sullivan & Augé, 2006

<==o †Glyptosaurinae
   |-- †Odaxosaurus
   `--+--+-- †Xestops
      |  `-- †Proxestops
      `--+--+-- †Paraplacosauriops
         |  `-- †Peltosaurus
         `--+--o †Melanosaurini
            |  |-- †Arpadosaurus
            |  `-- †Melanosaurus
            `--o †Glyptosaurini
               |-- †Placosaurus rugosus
               |-- †Glyptosaurus
               `--+-- †Proglyptosaurus huerfanensis
                  `--+-- †Paraglyptosaurus
                     `--+-- †Helodermoides
                        `--+-- †Placosaurus mongoliensis
                           `-- †Placosaurus estesi

<==o †Glyptosaurinae
   |-- †Odaxosaurus
   `--+-- Glyptosaurinae sp. et gen. indet.
      `--+-- †Proxestops
         `--+?- †“Xestops”
            |?- †Peltosaurus
            |--+-- †Arpadosaurus
            |  `-- †Melanosaurus
            `--+--+-- †Helodermoides
               |  `-- †Placosaurus
               `--+-- †Eoglyptosaurus
                  `--+-- †Glyptosaurus
                     `-- †Paraglyptosaurus

Reference(s):

• Augé, M., 2003: La faune de Lacertilia (Reptilia, Squamata) de l'Éocène inférieur de Prémontré (Bassin de Paris, France)
  [The lizard fauna (Reptilia, Squamata) from the lower Eocene locality of Prémontré (Paris Basin, France)].
  –Geodiversitas: Vol. 25, #3, pp. 539-574
• Augé, M. & Sullivan, R. M., 2006: A new genus, Paraplacosauriops (Squamata, Anguidae, Glyptosaurinae), from the Eocene of France.
  –Journal of Vertebrate Paleontology: Vol. 26, #1, pp. 133-137
• Carroll, R. L., 1988: Appendix. 594-648.
  in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
  –W. H. Freeman and company, New York, 1988, 698
• Conrad, J. L. & Norell, M. A., 2008: The braincases of two glyptosaurines (anguidae, Squamata) and anguid phylogeny.
  –American Museum Novitates: #3613, pp. 1-24 [doi: 10.1206/586.1]
• Sullivan, R. M., 1979: Revision of the Paleogene genus Glyptosaurus (Reptilia, Anguidae).
  –Bulletin of the American Museum of Natural History: #163, pp. 1-72
• Sullivan, R. M., 1986: The skull of Glyptosaurus sylvestris Marsh, 1871 (Lacertilia: Anguidae).
  –Journal of Vertebrate Paleontology: Vol. 6, #1, pp. 28-37
• Sullivan, R. M., 1989: Proglyptosaurus huerfanensis, new genus, new species: Glyptosaurine lizard (Squamata. Anguidae) from the Early Eocene of Colorado.
  –American Museum Novitates: #2949, pp. 1-8
• Sullivan, R. M. & Augé, M., 2006: Redescription of the holotype of Placosaurus rugosus Gervais 1848-1852 (Squamata, Anguidae, Glyptosaurinae) from the Eocene of France and a revision of the genus.
  –Journal of Vertebrate Paleontology: Vol. 26, #1, pp. 127-132
• Zonneveld, J.-P., Gunnell, G. F. & Bartels, W. S., 2000: Early Eocene fossil vertebrates from the southwestern Green River basin, Lincoln and Uinta Counties, Wyoming.
  –Journal of Vertebrate Paleontology: Vol. 20, #2, pp. 369-386