After Olson, 1985, and Sibley & Monroe, 1990
<==o Strigidae (pöllöt) [Recent: 22 genera /164 species] |?-o †Ornimegalonyx (Kuubanjättipöllöt) | |-- †O. oteroi | `-- †O. sp. |?-o †Grallistrix (Stilt-owls) | |-- †G. auceps (Kaua'i Stilt-owl) | |-- †G. erdmani (Maui Stilt-owl) | |-- †G. geleches (Moloka'i Stilt-owl) | `-- †G. orion (O'ahu Stilt-owl) |--o Striginae | |--+-- Otus (pöllöset) | | `-- Megascops Kaup, 1848 [Otus Pennant, 1769, in partim] (Screech owls, kirkupöllöset) | |-- Pyrroglaux podargina (Hartlaub & Finsch, 1872) Yamashina, 1938 [Otus podarginus] (palaunpöllönen) | |--o Margarobyas lawrencii (PL Sclater & Salvin, 1868) Olson & Suarez, 2008 [Otus lawrencii] (kuubanpöllönen) | | |-- M. l. exsul (Bangs, 1913) | | `-- M. l. lawrencii (PL Sclater & Salvin, 1868) | |--o Ptilopsis Kaup, 1848 | | |-- P. leucotis (Temminck, 1820) [Otus leucotis] (valkonaamapöllönen) | | `-- P. erlangeri (Ogilvie-Grant, 1906) | |-- Mimizuku gurneyi (Tweeddale, 1879) Hachisuka, 1934 (pisarapöllönen) | |-- Bubo Dumeril, 1805 [incl. Nyctea] (eagle owls; huuhkajat) | |--o Ketupa (kalahuuhkajat) | | |-- K. blakistoni (äyriäishuuhkaja) | | |-- K. zeylonensis (kalahuuhkaja) | | |-- K. flavipes (vuorikalahuuhkaja) | | `-- K. ketupu (viirukalahuuhkaja) | |--o Scotopelia (kalapöllöt) | | |-- S. peli (kalapöllö) | | |-- S. ussheri (ruskokalapöllö) | | `-- S. bouvieri (viirukalapöllö) | |-- Strix Linnaeus, 1758 (Viirupöllöt) | |-- Jubula lettii (Buttikofer, 1889) Bates, 1929 (takkupöllö) | |--o Lophostrix cristata (Daudin, 1800) Lesson, 1836 (töyhtöpöllö) | | |-- L. c. cristata (Daudin, 1800) | | |-- L. c. stricklandi PL Sclater & Salvin, 1859 | | `-- L. c. wedeli Griscom, 1932 | `--o Pulsatrix Kaup, 1848 (kultapöllöt) | |--o P. perspicillata (Latham, 1790) (kultapöllö) | | |-- P. p. boliviana L Kelso, 1933 | | |-- P. p. chapmani Griscom, 1932 | | |-- P. p. perspicillata (Latham, 1790) | | |-- P. p. pulsatrix (Wied-Neuwied, 1820) | | |-- P. p. saturata Ridgway, 1914 | | `-- P. p. trinitatis Bangs & TE Penard, 1918 | |-- P. koeniswaldiana (M Bertoni & W Bertoni, 1901) (kultakulmapöllö) | `--o P. melanota (Tschudi, 1844) (andienkultapöllö) | |-- P. m. melanota (Tschudi, 1844) | `-- P. m. philoscia Todd, 1947 |--o Surniinae | |--o Surnia ulula (Linnaeus, 1758) Dumeril, 1805 (hiiripöllö) | | |-- S. u. caparoch (Statius Muller, 1776) | | |-- S. u. tianschanica Smallbones, 1906 | | `-- S. u. ulula (Linnaeus, 1758) | |-- Glaucidium F Boie, 1826 (varpuspöllöt) | |-- Xenoglaux loweryi O'Neill & GR Graves, 1977 (partapöllö) | |--o Micrathene whitneyi (JG Cooper, 1861) Coues, 1866 (kaktuspöllö) | | |-- M. w. graysoni Ridgway, 1886 | | |-- M. w. idonea (Ridgway, 1914) | | |-- M. w. sanfordi (Ridgway, 1914) | | `-- M. w. whitneyi (Cooper,JG, 1861) | |-- Athene F Boie, 1822 [incl. Speotyto] (minervanpöllöt) | |-- Heteroglaux blewitti Hume, 1873 [Athene blewitti] (hindustaninpöllö) | |--o Aegolius Kaup, 1829 (helmipöllöt) | | |--o A. funereus (Linnaeus, 1758) (helmipöllö) | | | |-- A. f. beickianus Stresemann 1928 | | | |-- A. f. caucasicus (Buturlin, 1907) | | | |-- A. f. funereus (Linnaeus, 1758) | | | |-- A. f. magnus (Buturlin, 1907) | | | |-- A. f. pallens (Schalow, 1908) | | | |-- A. f. richardsoni (Bonaparte, 1838) | | | `-- A. f. sibiricus (Buturlin, 1910) | | |--o A. harrisii (Cassin, 1849) (keltahelmipöllö) | | | |-- A. h. dabbenei Olrog, 1979 | | | |-- A. h. harrisii (Cassin, 1849) | | | `-- A. h. iheringi (Sharpe, 1899) | | `--+--o A. acadicus (Gmelin, 1788) (amerikanhelmipöllö) | | | |-- A. a. acadicus (Gmelin, 1788) | | | `-- A. a. brooksi (JH Fleming, 1916 Auk 33 p.422) | | `--o A. ridgwayi (Alfaro, 1905) (ruskohelmipöllö) | | |-- A. r. ridgwayi (Alfaro, 1905) | | |-- A. r. rostratus (Griscom, 1930) | | `-- A. r. tacanensis RT Moore, 1947 | |-- Uroglaux dimorpha (Salvadori, 1874) Mayr, 1937 (papuanpöllö) | |--o †Sceloglaux albifacies (GR Gray, 1844) Kaup, 1848 (Whekau; Laughing Owl, White-faced Owl; kirkupöllö) [extinct since 1916?] | | |-- †S. a. albifacies (GR Gray, 1844) (South Island Laughing Owl) | | `-- †S. a. rufifacies Buller, 1904 (North Island Laughing Owl) | `--+?-o †Mascarenotus | | |-- †M. grucheti Mourer-Chauviré, Bour, Moutou & Ribes, 1994 (Réunion Owl) | | |-- †M. murivorus (Milne-Edwards, 1873) [Strix murivora Milne-Edwards, 1873; Carine murivora Günther & E Newton, 1879; Bubo leguati Rothschild, 1907 Athene murivora Rothschild, 1907] (Rodrigues Owl, Leguat's Owl; rodriguesinpöllö) [extinct since 1730's?] | | `-- †M. sauzieri (Newton & Gadow, 1893) [Strix sauzieri Newton & Gadow, 1893; Scops commersoni Oustalet, 1896; Strix newtoni Rothschild, 1907; Tyto sauzieri Hachisuka, 1953; Tyto newtoni Hachisuka, 1953] (Mauritius Owl, Commerson's Owl, Sauzier's Owl, Newton's Owl; mauritiuksenpöllö) [extinct since 1850's] | `-- Ninox Hodgson, 1837 (haukkapöllöt) `--o Asioninae |--o Pseudoscops Kaup, 1848 | |-- P. grammicus (Gosse, 1847) (jamaikanpöllö) | `--o P. clamator (Vieillot, 1808) [Asio clamator] (amerikansarvipöllö) | |-- P. c. clamator (Vieillot, 1808) | |-- P. c. forbesi (Lowery & Dalquest, 1951) | |-- P. c. midas (Schlegel, 1862) | `-- P. c. oberi (EH Kelso, 1936) |-- Nesasio solomonensis (Hartert, 1901) JL Peters, 1937 (isojalkapöllö) `-- Asio Brisson, 1760 (sarvipöllöt)
Reference(s):
- Lokki, J., Jännes, H. & Nikander, P. J., 1992: Otavan lintutieto.
–Kustannusosakeyhtiö Otava, Helsinki, Finland, 1992, pp. 1-440
Translated from: Perrins, C. M. (ed.), 1990: The Illustrated encyclopaedia of birds.
–Headline Book Publishing Plc., London, England, 1990 - Olson, S. L., 1985: The fossil record of birds. 79-239.
in Farner, D., King, J. & Parkes, K. (eds.), 1985: Avian Biology. vol VIII.
–Academic Press, New York. 1985. - Olson, S. L. & Hilgartner, W. B., 1982: Fossil and subfossil birds from the Bahamas. 22-57
in Olson, S. L., (ed.), 1982: The Vertebrates from the Bahamas.
–Smithsonian Contributions to Paleobiology: No. 48, pp. iv-65 - Sibley, C. G. & Monroe, B, L., 1990: Distribution and taxonomy of birds of the world.
–Yale University Press, New Haven & London, 1990. xxiv-1111.