After Sibley & Monroe, 1990, and Peterson, 2002-2011
<==o Megascops Kaup, 1848 [Otus Pennant, 1769, in partim] (Screech owls, kirkupöllöset) |-- M. flammeolus (Kaup, 1852) (ponderosapöllönen) |--o M. asio (Linnaeus, 1758) (idänkirkupöllönen) | |-- M. a. asio (Linnaeus, 1758) | |-- M. a. floridanus (Ridgway, 1874) | |-- M. a. hasbroucki (Ridgway, 1914) | |-- M. a. maxwelliae (Ridgway, 1877) | `-- M. a. mccallii (Cassin, 1854) |--+--o M. kennicottii (Elliot, 1867) (lännenkirkupöllönen) | | |-- M. k. aikeni Brewster, 1891 | | |-- M. k. bendirei (Brewster, 1882) | | |-- M. k. cardonensis (Huey, 1926) | | |-- M. k. kennicottii (Elliot, 1867) (lännenkirkupöllönen) | | |-- M. k. macfarlanei Brewster, 1891 | | |-- M. k. suttoni (RT Moore, 1941) | | |-- M. k. vinaceus Brewster, 1888 (Vinaceous Screech Owl) | | |-- M. k. xantusi Brewster, 1902 | | `-- M. k. yumanensis (AH Miller & L Miller, 1951) | |-- M. seductus (RT Moore, 1941) [Otus kennicottii seductus; Megascops kennicottii seductus] (meksikonkirkupöllönen) | `--o M. cooperi (Ridgway, 1878) [Otus kennicottii cooperi; Megascops kennicottii cooperi] (tyynenmerenpöllönen) | |-- M. c. cooperi (Ridgway, 1878) | `-- M. c. lambi (RT Moore & JT Marshall jr., 1959) [Otus kennicottii lambi; Megascops kennicottii lambi] (Oaxaca Screech Owl) |--o M. trichopsis (Wagler, 1832) (etelänkirkupöllönen) | |-- M. t. aspersus Brewster, 1888 | |-- M. t. mesamericanus (Van Rossem, 1932) | `-- M. t. trichopsis (Wagler, 1832) |--o M. choliba (Vieillot, 1817) (pampapöllönen) | |-- M. c. choliba (Vieillot, 1817) | |-- M. c. cruciger (Spix, 1824) | |-- M. c. decussatus (Lichtenstein, 1823) | |-- M. c. duidae (Chapman, 1929) | |-- M. c. luctisonus (Bangs & TE Penard, 1921) | |-- M. c. margaritae (Cory, 1915) | |-- M. c. surutus (L Kelso, 1941) | |-- M. c. uruguaiensis (Hekstra, 1982) | `-- M. c. wetmorei (Brodkorb, 1937) |-- M. koepckeae (Hekstra, 1982) (bolivianpöllönen) |--o M. roboratus (Bangs & Noble, 1918) (perunpöllönen) | |-- M. r. pacificus (Hekstra, 1982) (Coastal Screech Owl) | `-- M. r. roboratus (Bangs & Noble, 1918) (West Peruvian Screech Owl) |-- M. clarkii (L Kelso & EH Kelso, 1935) (panamanpöllönen) |-- M. barbarus (PL Sclater & Salvin, 1868) (guatemalanpöllönen) |--+-- M. petersoni (Fitzpatrick & O'Neill, 1986) [Otus marshalli petersoni; M. marshalli petersoni] (tumbesinpöllönen) | `-- M. marshalli (Weske & Terborgh, 1981) [Otus marshalli marshalli; M. marshalli marshalli] (pilvimetsäpöllönen) |--+-- M. colombianus (Traylor, 1952) [Otus ingens colombianus; M. ingens colombianus] (kolumbianpöllönen) | `--o M. ingens (Salvin, 1897) (andienpöllönen) | |-- M. i. venezuelanus (Phelps & Phelps, 1954) (venezuelanpöllönen) | `-- M. i. ingens (Salvin, 1897) (andienpöllönen) |--o M. watsonii (Cassin, 1849) (amazonianpöllönen) | |-- M. w. watsonii (Cassin, 1849) (Tawny-bellied Screech Owl) | `-- M. w. usta (PL Sclater, 1858) (Austral Screech Owl) |--+--o M. guatemalae (Sharpe, 1875) [Otus atricapillus guatemalae; M. atricapillus guatemalae] (Variable Screech Owl) | | |-- M. g. cassini (Ridgway, 1878) | | |-- M. g. dacrysistactus (RT Moore & JL Peters, 1939) | | |-- M. g. fuscus (RT Moore & JL Peters, 1939) | | |-- M. g. guatemalae (Sharpe, 1875) | | |-- M. g. hastatus Ridgway, 1887 | | |-- M. g. thompsoni (Cole, 1906) | | `-- M. g. tomlini (RT Moore, 1937) | |-- M. atricapilla (Temminck) 1822 [Otus atricapillus atricapillus; M. atricapilla atricapilla] (Black-capped Screech Owl; mustalakkipöllönen) | `-- M. hoyi (C Koenig & Straneck, 1989) [Otus atricapillus hoyi; M. atricapilla hoyi] (Hoy's Screech Owl) |-- M. vermiculatus Ridgway, 1887 (amerikanpöllönen) |-- M. roraimae (Salvin, 1897) (roraimanpöllönen) |-- M. centralis (Hekstra, 1982) |-- M. napensis (Chapman, 1928) |-- M. sanctaecatarinae (Salvin, 1897) (tupsupöllönen) |--o M. nudipes (Daudin, 1800) (puertoriconpöllönen) | |-- M. n. newtoni (Lawrence, 1860) | `-- M. n. nudipes (Daudin, 1800) `--o M. albogularis (Cassin, 1849) (valkokurkkupöllönen) |-- M. a. albogularis (Cassin, 1849) |-- M. a. macabrum (Bonaparte, 1850) |-- M. a. meridensis (Chapman, 1923) |-- M. a. obscurus (Phelps & WH Phelps jr., 1953) `-- M. a. remotus (Bond & Meyer de Schauensee, 1941)
Reference(s):
- Lokki, J., Jännes, H. & Nikander, P. J., 1992: Otavan lintutieto.
–Kustannusosakeyhtiö Otava, Helsinki, Finland, 1992, pp. 1-440
Translated from: Perrins, C. M. (ed.), 1990: The Illustrated encyclopaedia of birds.
–Headline Book Publishing Plc., London, England, 1990 - Olson, S. L., 1985: The fossil record of birds. 79-239.
in Farner, D., King, J. & Parkes, K. (eds.), 1985: Avian Biology. vol VIII.
–Academic Press, New York. 1985. - Peterson, A. P., 2002-2011: Zoonomen Nomenclatural data of Birds. (Updated: 2011-10-16)
–iNet: Zoonomen Nomenclatural data: http://www.zoonomen.net; [visited: 2011-12-19] - Sibley, C. G. & Monroe, B, L., 1990: Distribution and taxonomy of birds of the world.
–Yale University Press, New Haven & London, 1990. xxiv-1111