After Faivovich et al., 2005, and Frost et al., 2006, with Recent species from Frost, 2000-2002 [“AMNH Amphibian species of the World”]
<==o Hyla Laurenti, 1768 [Calamita Schneider, 1799; Hylaria Rafinesque, 1814; Hyas Wagler, 1830 (non Hyas Leach, 1815); Dendrohyas Wagler, 1830; Dryophytes Fitzinger, 1843; Epedaphus Cope, 1885] |--o H. arborea Group | |-- H. annectans (Jerdon, 1870) | |-- H. chinensis Günther, 1858 | |-- H. hallowellii Thomson, 1912 | |-- H. immaculata Boettger, 1888 | |-- H. intermedia Boulenger, 1882 | |-- H. meridionalis Boettger, 1874 (välimerenlehtismmakko) | |-- H. sanchiangensis Pope, 1929 | |-- H. sarda (De Betta, 1853) | |-- H. simplex Boettger, 1901 | |-- H. tsinlingensis Liu & Hu, 1966 | |-- H. ussuriensis Nikolsky, 1918 | |-- H. zhaopingensis Tang & Zhang, 1984 | `--+-- H. arborea (Linnaeus, 1758) [Rana arborea Linnaeus, 1758; Hyla viridis Laurenti, 1768] (euroopanlehtisammakko) | `-- H. savignyi Audouin, 1827 `--+--o H. cinerea Group | |-- H. squirella Bosc, 1800 | `--+-- H. cinerea (Schneider, 1799) | `-- H. gratiosa LeConte, 1857 “1856” `--+-- H. femoralis Bosc, 1800 |--o H. versicolor Group | |-- H. avivoca Viosca, 1928 | |-- H. chrysoscelis Cope, 1880 | `-- H. versicolor LeConte, 1825 `--o H. eximia Group |-- H. andersonii Baird, 1854 |-- H. arboricola Taylor, 1941 |-- H. bocourti (Mocquard, 1889) |-- H. plicata Brocchi, 1877 |-- H. suweonensisKuramoto, 1980 |-- H. wrightorum Taylor, 1939 `--+--+-- H. japonica Günther, 1859 “1858” | `-- H. walkeri Stuart, 1954 `--+-- H. arenicolor Cope, 1886 `--+-- H. euphorbiacea Günther, 1859 `-- H. eximia Baird, 1854
Reference(s):
- Faivovich, J., Haddad, C. F. B., García, P. C. A., Frost, D. R., Campbell, J. A. & Wheeler, W. C., 2005: Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic analysis.
–Bulletin of the American Museum of Natural History: #294, pp. 1-240 - Frost, D. R., 2000: Amphibian species of the World: An Online Reference.
–iNet: American Museum of Natural History, Department of Herpetology: http://research.amnh.org/herpetology/amphibia/index.html