After Faivovich et al., 2005, and Frost et al., 2006, with Recent species from Frost, 2000-2002 [“AMNH Amphibian species of the World”]
<==o Holoscirtus Peters, 1882 |--o H. bogotensis Group | |-- H. albopunctulatus (Boulenger, 1882) [Hyla albopunctulata] | |-- H. alytolylax (Duellmann, 1972) [Hyla alytolylax] | |-- H. bogotensis Peters, 1882 [Hyla bogotensis] | |-- H. callipeza (Duellmann, 1989) [Hyla callipeza] | |-- H. colymba (Dunn, 1931) [Hyla colymba] | |-- H. denticulentus (Duellmann, 1972) [Hyla denticulenta] | |-- H. estevesi (Rivero, 1968) | |-- H. jahni (Rivero, 1961) [Hyla jahni] | |-- H. lascinius (Rivero, 1969) [Hyla lascinia] | |-- H. lynchi (Ruiz-Carranza & Ardila-Robayo, 1991) [Hyla lynchi] | |-- H. palmeri (Boulenger, 1908) [Hyla palmeri] | |-- H. phyllognathus (Melin, 1941) [Hyla phyllognatha] | |-- H. piceigularis (Ruiz-Carranza & Lynch, 1982) [Hyla piceigularis] | |-- H. platydactylus (Boulenger, 1905) [Hyla platydactyla] | |-- H. simmonsi (Duellman, 1989) [H. simmonsi] | `-- H. torrenticola (Duellman & Altig, 1978) [Hyla torrenticola] `--+--o H. armatus Group | |-- H. armatus (Boulenger, 1902) [Hyla armata] | `-- H. charazani (Vellard, 1970) [Hyla charazani] `--o H. larinopygion Group |?- H. caucanus (Ardila-Robayo, Ruiz-Carranza & Roa-Trujillo, 1993) [Hyla caucana] |?- H. larinopygion (Duellman, 1973) [Hyla larinopygion] |?- H. lindae (Duellman & Altig, 1978) [Hyla lindae] |?- H. psarolaimus (Duellman & Hillis, 1990 [Hyla psarolaima] |?- H. ptychodactylus (Duellman & Hillis, 1990 [Hyla ptychodactyla] |?- H. sarampiona (Ruiz-Carranza & Lynch, 1982) [Hyla sarampiona] |?- H. staufferorum (Duellman & Coloma, 1993) [Hyla staufferorum] |-- H. tapichalaca (Kizirian, Coloma & Paredes-Recalde, 2003) [Hyla tapichalaca] `--+-- H. pacha (Duellman & Hillis, 1990) [Hyla pacha] `-- H. pantostictus (Duellman & Berger, 1982) [Hyla pantosticta]
Reference(s):
- Faivovich, J., Haddad, C. F. B., García, P. C. A., Frost, D. R., Campbell, J. A. & Wheeler, W. C., 2005: Systematic review of the frog family Hylidae, with special reference to Hylinae: phylogenetic analysis and taxonomic analysis.
–Bulletin of the American Museum of Natural History: #294, pp. 1-240 - Frost, D. R., 2000: Amphibian species of the World: An Online Reference.
–iNet: American Museum of Natural History, Department of Herpetology: http://research.amnh.org/herpetology/amphibia/index.html