After Tyler, Jerzma´nska, Bannikov & ´Swidnicki, 1993, and Nelson, 1994, with Recent species from Froese & Pauly, 2001 [“The Fishbase”]
<==o Triacanthodidae sensu Tyler, 1980 (vaarnakalat; spikefishes) [Nelson: 11/20] [Fishbase: 11/21] |--o Hollardiinae Tyler, 1968 [2/5] | |-- †Prohollardia avita Tyler et al., 1993, Olig. Eu. | `--+--o Hollardia [3] | | |-- H. goslinei Tyler, 1968 | | |-- H. hollardi Poey, 1861 (Reticulate spikefish) | | `-- H. meadi Tyler, 1966 (Spotted spikefish) | `--o Parahollardia [2] | |-- P. lineata (Longley, 1935) (Jambeau) | `-- P. schmidti Woods, 1959 `--o Triacanthodinae Tyler, 1968 [Nelson: 9/15] [Fishbase: 9/16] |-- †Carpathospinosus propheticus Tyler et al., 1993, Olig. Eu. |--o Atrophacanthus [1 Recent species] | |-- †A. serratus | |-- †A. collettei | |?- †A. trispinosus [Cephalacanthus] | `-- A. japonicus (Kamohara, 1941) |--o Bathyphylax [2] | |-- B. bombifrons Myers, 1934 | `-- B. omen Tyler, 1966 |--o Halimochirurgus [2] | |-- H. alcocki Weber, 1913 | `-- H. centriscoides Alcock, 1899 (Longsnout spikefish) |-- Johnsonina eriomma Myers, 1934 |--o Macrorhamphosodes [2] | |-- M. platycheilus Fowler, 1934 (Trumpetsnout spikefish) | `-- M. uradoi (Kamohara, 1933) (Trumpetsnout) |-- Mephisto fraserbrunneri Tyler, 1966 |--o Paratriacanthodes [2] | |-- P. herrei Myers, 1934 | `-- P. retrospinis Fowler, 1934 (Sawspine spikefish) |--o Triacanthodes [4] | |-- T. anomalus (Temminck & Schlegel, 1850) | |-- T. ethiops Alcock, 1894 (Shortsnout spikefish) | |-- T. indicus Matsuura, 1982 | `-- T. intermedius Matsuura & Fourmanoir, 1984 `-- Tydemania navigatoris Weber, 1913 (Fleshy-lipped spikefish)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Frickhinger, K. A., 1995: Fossil Atlas – Fishes.
–Mergus – Publishers for Natural History and Pet Books, Hans A. Baensch, Malle, Germany, 1-1088 - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 5: Kalat, sammakkoeläimet, matelijat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beauté Du Monde Animal: IX Reptiles/Amphibiens & X Poissons.
–Rizzoli Editore, Milano, 1968 - Nelson, J. S., 1994: Fishes of the world.
–John Wiley & Sons Inc., New York, 1994, xx-600 - Tyler, J. C., Jerzma´nska, A., Bannikov, A. F. & ´Swidnicki, J., 1993: Two new genera and species of Oligocene spikefishes (Tetraodontiformes: Triacanthonidae), the first fossils of the Hollardiinae and Triacanthodidae.
–Smithsonian Contributions to Paleobiology: No. 75, pp. vi-27