After Nelson, 1994, with Recent species from Froese & Pauly, 2001 [“The Fishbase”]
<==o Psychrolutidae (pallosimput) [N: 7/29] [FB: 10/30] |?-o Ambophthalmos [3] [=? “Neophrynichthys” [Nelson, 1994]] | |-- A. angustus (Nelson, 1977) Pale toadfish | |-- A. eurystigmatephoros Jackson & Nelson, 1999 | `-- A. magnicirrus (Nelson, 1977) |?- Thecopterus aleuticus Smith, 1904 |--o Cottunculinae [N: 5/12] | |--o Cottunculus [6] | | |-- C. granulosus Karrer, 1968 | | |-- C. microps Collett, 1875 (Polar sculpin) | | |-- C. nudus Nelson, 1989 (Bonyskull toadfish) | | |-- C. sadko Essipov, 1937 | | |-- C. spinosus Gilchrist, 1906 | | `-- C. thomsonii (Günther, 1882) (Pallid sculpin) | |-- Dasycottus setiger Bean, 1890 (Spinyhead sculpin) | |-- Eurymen gyrinus Gilbert & Burke, 1912 (Smooth-cheek sculpin) | |--o Malacocottus [3] | | |-- M. gibber Sakamoto, 1930 | | |-- M. kincaidi Gilbert & Thompson, 1905 (Blackfin sculpin) | | `-- M. zonurus Bean, 1890 (Darkfin sculpin) | `?- “Neophrynichthys” [2] [Nelson, 1994] `--o Psychrolutinae [3/17] |--o Ebinania [3] | |-- E. brephocephala (Jordan & Starks, 1903) | |-- E. costaecanariae (Cervigón, 1961) | `-- E. vermiculata Sakamoto, 1932 |-- Neophrynichthys latus (Hutton, 1875) (Dark toadfish) `--o Psychrolutes [Cottunculoides, Gilbertidia] [11] |-- P. inermis (Vaillant, 1888) |-- P. macrocephalus (Gilchrist, 1904) |-- P. marcidus (McCulloch, 1926) (Blobfish) |-- P. marmoratus (Gill, 1888) |-- P. occidentalis Fricke, 1990 (Western Australian sculpin) |-- P. paradoxus Günther, 1861 (Tadpole sculpin) |-- P. phrictus Stein & Bond, 1978 (Blob sculpin) |-- P. pustulosus (Schmidt, 1937) |-- P. sigalutes (Jordan & Starks, 1895) (Soft sculpin) |-- P. sio Nelson, 1980 `-- P. subspinosus (Jensen, 1902)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 5: Kalat, sammakkoeläimet, matelijat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beauté Du Monde Animal: IX Reptiles/Amphibiens & X Poissons.
–Rizzoli Editore, Milano, 1968 - Nelson, J. S., 1994: Fishes of the world.
–John Wiley & Sons Inc., New York, 1994, xx-600