After Froese & Pauly, 2001 [“The Fishbase”]
<==o Cottus Linnaeus, 1758; R. As. Mioc. NZ. L. Olig. R. Eu. Mioc. Plioc. R. NA. (sculpins; kivisimput) [N: 34] [FB: 45] |-- †C. brevis |-- C. aleuticus Gilbert, 1896 (Coastrange sculpin) |-- C. amblystomopsis Schmidt, 1904 (Sakhalin sculpin) |-- C. asper Richardson, 1836 (Prickly sculpin) |-- C. asperrimus Rutter, 1908 (Rough sculpin) |--o C. bairdii -group | |-- C. baileyi Robins, 1961 (Black sculpin) | |--+-- C. caeruleomentum Kinziger, Raesly & Neely, 2000 (Blue Ridge sculpin) | | `--o C. bairdii Girard, 1850 (Mottled sculpin) | | |-- C. b. bairdii Girard, 1850 | | |-- C. b. punctulatus | | |-- C. b. semiscaber | | `-- C. b. kumlieni | |-- C. cognatus Richardson, 1836 (Slimy sculpin) | |-- C. hypselurus Robins & Robison, 1985 (Ozark sculpin) | `-- C. paulus Williams, 2000 (Pygmy sculpin) |-- C. beldingii Eigenmann & Eigenmann, 1891 (Paiute sculpin) |--o C. carolinae -group | `-- C. carolinae (Gill, 1861) (Banded sculpin) |-- C. confusus Bailey & Bond, 1963 (Shorthead sculpin) |-- C. czerskii Berg, 1913 (Cherskii's sculpin) |-- C. echinatus Bailey & Bond, 1963 (Utah Lake sculpin) |-- C. extensus Bailey & Bond, 1963 (Bear Lake sculpin) |-- C. ferrugineus Heckel & Kner, 1858 |-- C. girardi Robins, 1961 (Potomac sculpin) |--o C. gobio Linnaeus, 1758 (kivisimppu; Bullhead) | |-- C. g. gobio Linnaeus, 1758 (kivisimppu; Bullhead) | `-- C. g. jaxartensis Berg, 1916 |-- C. greenei (Gilbert & Culver, 1898) (Shoshone sculpin) |-- C. gulosus (Girard, 1854) (Riffle sculpin) |-- C. hangiongensis Mori, 1930 |-- C. kaganowskii Berg, 1932 (Anadyr sculpin) |-- C. kazika Jordan & Starks, 1904 |--o C. kesslerii Dybowski, 1874 (Kessler's sculpin) | |-- C. k. bauntovi Taliev, 1946 (Baunt sand sculpin) | `-- C. k. kesslerii Dybowski, 1874 (Kessler's sculpin) |--o C. klamathensis Gilbert, 1898 (Marbled sculpin) | |-- C. k. polyporus | `--+-- C. k. macrops | `-- C. k. klamathensis Gilbert, 1898 |-- C. leiopomus Gilbert & Evermann, 1894 (Wood River sculpin) |-- C. marginatus (Bean, 1881) (Margined sculpin) |-- C. nasalis Berg, 1933 (Tubenose sculpin) |-- C. nozawae Snyder, 1911 |-- C. perplexus Gilbert & Evermann, 1894 (Reticulate sculpin) |-- C. petiti Bacescu & Bacescu-Mester, 1964 |-- C. pitensis Bailey & Bond, 1963 (Pit sculpin) |--o C. poecilopus Heckel, 1837 (kirjoeväsimppu; Alpine bullhead) | |-- C. p. poecilopus Heckel, 1837 (Alpine bullhead) | `-- C. p. volki (Taranetz, 1933) |-- C. pollux Günther, 1873 (Japanese fluvial sculpin) |-- C. princeps Gilbert, 1898 (Klamath Lake sculpin) |-- C. reinii Hilgendorf, 1879 |-- C. rhotheus (Smith, 1882) (Torrent sculpin) |-- C. ricei (Nelson, 1876) (Spoonhead sculpin) |-- C. sibiricus Kessler, 1899 (Siberian sculpin) |-- C. spinulosus Kessler, 1872 (Turkestan sculpin) |-- C. tenuis (Evermann & Meek, 1898) (Slender sculpin) |-- C. hubbsi |-- C. bendirei `-- C. paulus
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Frickhinger, K. A., 1995: Fossil Atlas – Fishes.
–Mergus – Publishers for Natural History and Pet Books, Hans A.Baensch, Malle, Germany, 1995, pp. 1-1088 - Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org - Kinziger, A. P., Raesly, R. L. & Neely, D. A., 2000: New species of Cottus (Teleostei: Cottidae) from the middle Atlantic eastern United States.
–Copeia: Vol. 100, #4, pp. 1007-1018 - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 5: Kalat, sammakkoeläimet, matelijat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beauté Du Monde Animal: IX Reptiles/Amphibiens & X Poissons.
–Rizzoli Editore, Milano, 1968 - Nelson, J. S., 1994: Fishes of the world.
–John Wiley & Sons Inc., New York, 1994, xx-600