After Dyer & Chernoff, 1996, with Recent species from Froese & Pauly, 2001- [“The Fishbase”]

<==o Atherinidae Günther (silversides; hopeakyljet)
   |?- †Prosphyraena M. Eoc. Eu. 
   |?-o †Rhamphognathus [Mesogaster] M. Eoc. Eu.
   |  `-- †R. sphyraenoides
   |?- †Zanteclites Mioc. NA. 
   |-- Atherinomorinae Dyer & Chernoff, 1996
   `--+--o Craterocephalinae Dyer & Chernoff, 1996
      |  |?- Allanetta Whitley [not recognized by the Fishbase]
      |  `--+?- Quirichthys Whitley [not recognized by the Fishbase]
      |     `-- Craterocephalus McCulloch [25]
      `--o Atherininae
         |--o Atherinosoma Gastelnau [2]
         |  |-- A. elongata (Klunzinger, 1880)
         |  `-- A. microstoma (Günther, 1861) Small-mouth hardyhead
         `--+?- Atherinason hepsetoides (Richardson, 1843) Whitney [sedis mutabilis]
            |?-o Leptatherina Pavlov, Ivantsoff, Last, & Crowley, 1988 [2] [sedis mutabilis]
            |  |-- L. presbyteroides (Richardson, 1843)
            |  `-- L. wallacei (Prince, Ivantsoff & Potter, 1982) (Western hardyhead)
            `--+--o Kestratherina Pavlov, Ivantsoff, Last, & Crowley, 1988 [2]
               |  |-- K. brevirostris Pavlov, Ivantsoff, Last, & Crowley, 1988 (Short-snout hardyhead)
               |  `-- K. esox (Klunzinger, 1872)
               `--o Atherina Linnaeus, 1758; M. Eoc. R. Eu. Mioc. R. As. R. NA. (hopeakyljet) [5]
                  |-- A. boyeri Risso, 1810 (Big-scale sand smelt)
                  |-- A. breviceps Valenciennes, 1835 (Cape silverside)
                  |-- A. hepsetus Linnaeus, 1758 (Mediterranean sand smelt)
                  |-- A. lopeziana Rossignol & Blache, 1961
                  `-- A. presbyter Cuvier, 1829 (hopeakylki; Sand smelt)

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