After Carroll, 1988 and Nelson, 1994, with Recent species from Froese & Pauly, 2001 [“The Fishbase”]

<==o Albuloidei Greenwood et al., 1966 (naiskalat)
   |?-o †Phyllodontidae
   |  |?- †Casierius L. Cret. Eu. NA.
   |  |?- †Eodiaphyodus U. Cret. WAf.
   |  |--o †Egertonia Cocchi, 1866; U. Paleoc. L. Eoc. Eu., U. Eoc. L. Olig. NAf.
   |  |  |-- †E. isodonta Cocchi, 1866
   |  |  `-- †E. stromeri
   |  |-- †Paralbula U. Cret. Mioc. NA., L. Eoc. Eu., U. Eoc. NAf.
   |  |-- †Phyllodus L. Paleoc. U. Eoc. Eu., L. Paleoc. Mioc. NA.
   |  `-- †Pseudoegertonia L. Paleoc. WAf.
   |--o Pterothrissidae [Albulidae: Pterothrissinae]
   |  |-- †Hajulia multidens U. Cret. SWAs.
   |  |--o †Thrissipteroides U. Cret. Eu. SWAs.
   |  |  `-- †T. elongatus
   |  `--o Istieus U. Cret. Eu. SWAs. [Pterothrissus Hilgendorf, 1877; ?M. Jur. U. Jur. U. Cret. Eoc. Eu. Olig. Aus. R. Atl. Pac.] [2]
   |     |-- †I. macrocephalus
   |     |-- I. belloci (Cadenat, 1937) (Longfin bonefish)
   |     `-- I. gissu (Hilgendorf, 1877) [Pterothrissus gissu Hilgendorf, 1877] (Japanese gissu)
   `--o Albulidae Bleeker, 1859 (naiskalat; Bonefishes)
      |-- †Coriops U. Cret. NA. 
      |-- †Kleinpellia U. Cret. NA. 
      |--o †Lebonichthys U. Cret. SWAs.
      |  |-- †L. lewisi
      |  `-- †L. gracilis
      `--+-- †Deltaichthys albuloides Fielitz & Bardack, 1992
         `--+?- †Farinichthys gigas Gallo & de Figueiredo, 2002, Pal. NESA.
            `--o Albula ?U. Cret. ?NA. L.-M. Eoc. Eu. R. Oc.
               |?- A. glossodonta (Forsskål, 1775) (Roundjaw bonefish)
               |-- A. nemoptera (Fowler, 1911) (Threadfin bonefish)
               |?- A. neoguinaica Valenciennes, 1847 (Sharpjaw bonefish)
               `-- A. vulpes (Linnaeus, 1758) (Bonefish)

Note:

Nelson (1994) recognizes only two recent species of Albula, but Fishbase recognizes four, also Fishbase still uses genus-name Pterothrissus, which is is synonym of Istieus according to Carroll (1988).

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