After Sereno, 2000, Makovicky, 2001, Xu et al., 2002, Xu et al., 2006, Makovicky & Norell, 2006, and Farke, Maxwell, Cifelli & Wedel, 2014
Alternative phylogeny of Ceratopsia after Sereno, 2000, and You & Dodson, 2003
Alternative phylogeny of Ceratopsia after Makovicky, 2001
Alternative phylogeny of Ceratopsia after You & Dodson, 2003
Alternative phylogeny of Ceratopsia after Chinnery, 2004
<==o †Ceratopsia Marsh, 1890 (sarvinaamat; horned dinosaurs) |-- †Yinlong downsi Xu, Forster, Clark & Mo, 2006 `--+--o †Chaoyangsauridae Zhao, Cheng, Xu & Makovicky, 2006 | |-- †Chaoyangsaurus youngi Zhao, Cheng & Xu, 1999 | |-- †Hualianceratops wucaiwanensis Han, Forster, Clark & Xu, 2015 | `-- †Xuanhuaceratops niei Zhao, Cheng, Xu & Makovicky, 2006 `--+--o †Psittacosauria Osborn, 1923 | `--o †Psittacosauridae Osborn, 1923 | |?- †Hongshanosaurus houi You, Xu & Wang, 2003 | `--o †Psittacosaurus Osborn, 1923a [Protiguanodon Osborn, 1923a] sensu Sereno, 2010 | |?- †P. sattayaraki Buffetaut & Suteenthorn, 1992 [nomen dubium] | |?- †P. ordosensis Russell & Zhao, 1996; E.Cret. EAs. [=? P. sinensis] | |?- †P. mazongshanensis Xu, 1997 | |?- †P. xinjiangensis Sereno & Chao, 1988; E.Cret. EAs. | |-- †P. mongoliensis Osborn, 1923; E.Cret. EAs. [taxonomic history] | `--+--+?- †P. meileyingensis Sereno, Chao, Cheng & Rao, 1988; E.Cret. EAs. | | `--+?- †P. lujiatunensis Zhou, Gao, Fox & Chen, 2006; E.Cret. EAs. | | `?- †P. major Sereno, Zhao, Brown & Tan, 2007; E.Cret. EAs. | `--+-- †P. gobiensis Sereno, Zhao & Lin, 2009; E.Cret. EAs. | |-- †P. sibiricus Voronkevich & Averianov, 2000; E.Cret. EAs. | `--+?- †P. neimongoliensis Russell & Zhao, 1996; E.Cret. EAs. | `-- †P. sinensis Young, 1958a; E.Cret. EAs. `--o †Neoceratopsia Sereno, 1991 [Neoceratopia] |?- †Craspedodon lonzeensis Dollo, 1883 sensu Godefroit & Lambert, 2007 [tooth-taxa, could as well be an ornithopod] |-- †Liaoceratops yanzigouensis Xu, Makovicky, Wang, Norell & You, 2002 `--+-- †Aquilops americanus Farke, Maxwell, Cifelli & Wedel, 2014 `--+--+?- †Auroraceratops rugosus You, Li, Ji, Lamanna & Dodson, 2005 sensu Farke et al., 2014 | `-- †Yamaceratops dorngobiensis Makovicky & Norell, 2006 `--+--+?- †Helioceratops brachygnathus Jin, Chen, Zan & Godefroit, 2009 | `-- †Archaeoceratops oshimai Dong & Azuma, 1997 [“Archaeoceratopidae”] `--+-- †Koreaceratops hwaseongensis Lee, Ryan & Kobayashi, 2011 `--+--o †Leptoceratopsidae Nopcsa, 1923 | |-- †Asiaceratops salsopaludalis Nesov, Kaznyshkina & Cherepanov, 1989 [nomen dubium] | `--+-- †Cerasinops hodgskissi Chinnery & Horner, 2007 | `--+-- †Montanoceratops cerorhynchus (Brown & Schlaikjer, 1942) Sternberg, 1951 [Leptoceratops cerorhynchus Brown & Schlaikjer, 1942] | `--+-- †Prenoceratops pieganensis Chinnery, 2004 | `--+--+-- †Udanoceratops tschizhovi Kurzanov, 1992 | | `-- †Leptoceratops gracilis Brown, 1914c | `--+-- †Zhuchengceratops inexpectus Xu, Wang, Zhao, Sullivan & Chen, 2010 | `--+-- †Gryphoceratops morrisoni Ryan, Evans, Currie, Brown & Brinkman, 2012 | `-- †Unescoceratops koppelhusae Ryan, Evans, Currie, Brown & Brinkman, 2012 `--o †Coronosauria Sereno, 1986 |-- †Graciliceratops mongoliensis Sereno, 2000 `--+--o †Protoceratopidae Granger & Gregory, 1923 [incl.? Bagaceratopidae Alifanov, 2003] | |?- †Gobiceratops minutus Alifanov, 2008 | |?- †Lamaceratops tereschenkoi Alifanov, 2003 | |?- †Magnirostris dodsoni You & Dong, 2003 | |?- †Platyceratops tatarinovi Alifanov, 2003 | |--o †Protoceratops Granger & Gregory, 1923 | | |-- †P. andrewsi Granger & Gregory, 1923 | | `-- †P. hellenikorhinus Lambert, Godefroit, Li, Shang & Dong, 2001 | `--+-- †Ajkaceratops kozmai Ösi, Butler & Weishampel, 2010 | `-- †Bagaceratops rozhdestvenskyi Marya´nska & Osmólska, 1975 [Breviceratops kozlowskii (Marya´nska & Osmólska, 1975)] `--o †Ceratopsoidea Hay, 1902 [Ceratopoidea] `--o †Ceratopsomorpha |-- †Zuniceratops christopheri Wolfe & Kirkland, 1998 `--+-- †Turanoceratops tardabilis Nesov, Kaznyshkina & Cherepanov, 1989 `-- †Ceratopsidae Marsh, 1890a [Ceratopidae Marsh, 1890a emend. ??; Agathaumidae Cope]
Ceratopsia incertae sedis:
†Kulceratops kulensis Nessov, 1995
†Microceratops Bohlin, 1953 sensu Sereno, 2000 [nomen dubium]
†M. gobiensis Bohlin, 1953 [nomen dubium]
†M. sulcidens Bohlin, 1953 [nomen dubium]
†Notoceratops bonarellii Tapia, 1918 [nomen dubium]
†Serendipaceratops arthurcclarkei Rich & Vickers-Rich, 2003 [An Australian leptoceratopsid?]
After Han, Forster, Clark & Xu, 2015<==o †Ceratopsia |--o †Psittacosauria | |--o †Chaoyangsauridae | | |-- †Yinlong | | |-- †Chaoyangsaurus | | |-- †Hualianceratops | | `-- †Xuanhuaceratops | `--o †Psittacosauridae | `-- †Psittacosaurus `--o †Neoceratopsia |-- †Liaoceratops `--+-- †Aquilops |-- †Auroraceratops |-- †Archaeoceratops `--+-- †Yamaceratops `--+-- †Graciliceratops |-- †Asiaceratops |--o †Leptoceratopsidae | |-- †Cerasinops | `--+-- †Prenoceratops | |--+-- †Koreaceratops | | `-- †Montanaceratops [sic] | `--+-- †Unescoceratops | |-- †Zhuchengceratops | |-- †Udanoceratops | `-- †Leptoceratops `--o †Coronosauria |--o †Protoceratopidae | |-- †Protoceratops | `-- †Bagaceratops `--o †Ceratopsoidea `--o †Ceratopsomorpha |-- †Zuniceratops `-- †Ceratopsidae Reference(s):
|
After Farke, Maxwell, Cifelli & Wedel, 2014<==o †Ceratopsia |-- †Psittacosauria `--o †Neoceratopsia |-- †Liaoceratops `--+-- †Aquilops `--+--+-- †Auroraceratops | `-- †Yamaceratops `--+--+-- †Archaeoceratops | `-- †Helioceratops `--+-- †Koreaceratops `--+--o †Leptoceratopsidae | |-- †Asiaceratops | `--+-- †Cerasinops | `--+-- †Montanaceratops [sic] | `--+-- †Prenoceratops | `--+--+-- †Udanoceratops | | `-- †Leptoceratops | `--+-- †Zhuchengceratops | `--+-- †Gryphoceratops | `-- †Unescoceratops `--o †Coronosauria |-- †Graciliceratops `--+--o †Protoceratopidae | |-- †Protoceratops | `--+-- †Ajkaceratops | `-- †Bagaceratops `--o †Ceratopsoidea `--o †Ceratopsomorpha |-- †Zuniceratops `--+-- †Turanoceratops `-- †Ceratopsidae Reference(s):
|
After Chinnery, 2004a, b<==o †Ceratopsia |-- †Psittacosauria `--o †Neoceratopsia |--o †“Chaoyangsauridae” | |-- †Chaoyangsaurus | `-- †Asiaceratops `--+-- †Liaoceratops `--+-- †Archaeoceratops `--+--o †“Leptoceratopsidae” | |-- †Montanoceratops | `--+-- †Prenoceratops | `--+-- †Udanoceratops | `-- †Leptoceratops `--o †Coronosauria |--o †Protoceratopidae | |-- †Graciliceratops | |-- †Bagaceratops | `-- †Protoceratops `--o †Ceratopsoidea `--o †Ceratopsomorpha |-- †Zuniceratops `--+-- †Turanoceratops `-- †Ceratopsidae |
After You & Dodson, 2003<==o †Ceratopsia |-- †Psittacosauria `--o †Neoceratopsia |-- †Chaoyangsaurus `--+-- †Liaoceratops `--+-- †Archaeoceratops `--o †Coronosauria |--o †Protoceratopidae | |-- †Graciliceratops | |-- †Bagaceratops | `-- †Protoceratops `--o †Ceratopsoidea |--o †Leptoceratopsidae | |?- †Udanoceratops | |-- †Montanoceratops | `-- †Leptoceratops `--o †Ceratopsomorpha |-- †Zuniceratops `--+-- †Turanoceratops `-- †Ceratopsidae |
After Sereno, 2000, and You & Dodson, 2003<==o †Ceratopsia |-- †Psittacosauria `--o †Neoceratopsia |-- †Chaoyangsaurus `--+-- †Liaoceratops `--+-- †Archaeoceratops `--+--o †“Leptoceratopidae” | |?- †Udanoceratops | `-- †Leptoceratops `--o †Coronosauria |--o †Protoceratopidae | |-- †Graciliceratops | |-- †Bagaceratops | `-- †Protoceratops `--o †Ceratopsoidea |-- †Montanoceratops `--o †Ceratopsomorpha |-- †Zuniceratops `--+-- †Turanoceratops `-- †Ceratopsidae |
After Makovicky, 2001<==o †Ceratopsia |--+-- Ceratopsia genera et species indet. [China] | `-- †Chaoyangsaurus `--+-- †Psittacosauria `--o †Neoceratopsia |-- †Archaeoceratops `--+--o †“Leptoceratopidae” | |-- †Leptoceratops | |-- †Montanoceratops | `-- †Udanoceratops `--o †Coronosauria |--o †Protoceratopidae | |?- †Graciliceratops | |-- †Bagaceratops | `-- †Protoceratops `--o †Ceratopsoidea `--o †Ceratopsomorpha |-- †Zuniceratops `--+?- †Turanoceratops `-- †Ceratopsidae |
Reference(s):
- Chinnery, B. J., 2004: Description of Prenoceratops pieganensis gen. et sp. nov. (Dinosauria: Neoceratopsia) from the Two Medicine Formation of Montana.
–Journal of Vertebrate Paleontology: Vol. 24, #3, pp. 572-590 - Chinnery, B. J. & Weishampel, D. B., 1998: Montanoceratops cerorhynchus (Dinosauria: Ceratopsia) and relationships among basal neoceratopsians.
–Journal of Vertebrate Paleontology: Vol. 18, #3, pp. 369-585 - Dopson, P., 1996: The Horned Dinosaurs.
–Princeton University Press, Pinceton, New Jersey, pp. xiv-346 - Dopson, P., & Currie, P. J., 1990: Neoceratopsia. 593-618.
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Godefroit, P. & Lambert, O., 2007: A re-appraisal of Craspedodon lonzeensis Dollo, 1883 from the Upper Cretaceous of Belgium: the first record of a neoceratopsian dinosaur in Europe.
–Bulletin de l'Institut Royal des Sciences Naturelles de Belgique - Sciences de la Terra / Bulletin van het Koninkljik Belgisch Instituut voor Natuurwetenschappen - Aardwetenschappen: Vol. 77, pp. 83-93 - Farke, A. A., Maxwell, W. D., Cifelli, R. L. & Wedel, M. J., 2014: A Ceratopsian Dinosaur from the Lower Cretaceous of Western North America, and the Biogeography of Neoceratopsia.
–Plos One: Vol. 9, #12, art.nr. e112055 [doi: 10.1371/journal.pone.0112055] - Han, F., Forster, C. A., Clark, J. M. & Xu, X., 2015: A New Taxon of Basal Ceratopsian from China and the Early Evolution of Ceratopsia.
–Plos One: Vol. 10, #12, art.nr. e0143369 (pp. 1-23) [doi: 10.1371/journal.pone.0143369] - Jin, L., Chen, J., Zan, S. & Godefroit, P., 2009: A New Basal Neoceratopsian Dinosaur from the Middle Cretaceous of Jilin Province, China.
–Acta Geologica Sinica - English Edition: Vol. 83, #2, pp. 200-206 [doi: 10.1111/j.1755-6724.2009.00023.x] - Keesey, T. M., 1996-2000: The Dinosauricon
–iNet: http://dinosauricon.com/ - Lambert, O, Godefroit, P, Li, H, Shang, C.-Y. & Dong,Z.-M., 2001: A new species of Protoceratops (Dinosauria, Neoceratopsia) from the Late Cretaceous of Inner Mongolia (P. R. China).
–Bulletin de l'Institut Royal des Sciences Naturelles de Belgique - Science de la Terre: Vol. 71, suppl. 1, pp. 5–28. - Lee, Y.-N., Ryan, M. & Kobayashi, Y., 2011: The first ceratopsian dinosaur from South Korea.
–Naturwissenschaften: Vol. 98, #1, pp. 39-49 [doi: 10.1007/s00114-010-0739-y] - Makovicky, P. J. & Norell, M. A., 2006: Yamaceratops dorngobiensis, a new primitive ceratopsian (Dinosauria: Ornithischia) from the Cretaceous of Mongolia.
–American Museum Novitates: #3530, pp. 1-42 - Ösi, A., Butler, R. J. & Weishampel, D. B., 2010: A Late Cretaceous ceratopsian dinosaur from Europe with Asian affinities.
–Nature: Vol. 465, pp. 466-468 [doi: 10.1038/nature09019] - Ryan, M. J., Evans, D. C., Currie, P. J., Brown, C. M. & Brinkman, D., 2012: New leptoceratopsids from the Upper Cretaceous of Alberta, Canada.
–Cretaceous Research: Vol. 35, pp. 69-80 [doi: http://dx.doi.org/10.1016/j.cretres.2011.11.018] - Sereno, P. C., 2000: The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia. 480-516
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, xxxix-696 - Sereno, P. C., 1990: Psittacosauria. 579-592
in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.), 1990: The Dinosauria.
–University of California Press, Berkley, Los Angeles, Oxford, 1990 xvi-733 - Sereno, P. C., 2010: Taxonomy, cranial morphology, and relationships of parrot-beaked dinosaurs (Ceratopsia: Psittacosaurus). (21-58)
in Ryan, M. J., Chinnery-Allgeier, B. J. & Eberth, D. A. (eds.), 2010: New Perspectives on Horned Dinosaurs: The Royal Tyrrell Museum Ceratopsian Symposium. –Indiana University Press, Bloomington,Indianapolis, 2010, 656 p. - Sereno, P. C. & Chao, S., 1988: Psittacosaurus xinjiangensis (Ornithischia: Ceratopsia), a new psittacosaur from the Lower Cretaceous of Northwestern China.
–Journal of Vertebrate Paleontology: Vol. 8, #4, pp. 353-365 - Sereno, P. C., Chao, S., Cheng, Z. & Rao, C., 1988: Psittacosaurus meileyingensis (Ornithischia: Ceratopsia), a new psittacosaur from the Lower Cretaceous of Northeastern China.
–Journal of Vertebrate Paleontology: Vol. 8, #4, pp. 366-377 - Sereno, P. C., Zhao, X., Brown, L. & Tan, L., 2007: New psittacosaurid highlights skull enlargement in horned dinosaurs.
–Acta Palaeontologica Polonica: Vol. 52, #2, pp. 275-284 [http://app.pan.pl/acta52/app52-275.pdf] - Sereno, P. C., Zhao, X. & Lin, T., 2010: A new psittacosaur from Inner Mongolia and the parrot-like structure and function of the psittacosaur skull.
–Proceedings of the Royal Society of London: Biological series: Vol. 277, #1, pp. 199-209 [doi: 10.1098/rspb.2009.0691] - You, H., 2000: A new phylogeny of basal Neocetopsia.
–Journal of Vertebrate Paleontology: Vol. 20, #3, suppl. to #3, 80A - You, H.-l. & Dodson, P., 2003: Redescription of neoceratopsian dinosaur Archaeoceratops and early evolution of Neoceratopsia.
–Acta Palaeontologica Polonica: Vol. 48, #2, pp. 261-272 - You, H.-l., Li, D., Ji, Q., Lamanna, M. C. & Dodson, P., 2005: On a new genus of basal neoceratopsian dinosaur from the Early Cretaceous of Gansu Province, China.
–Acta Geologica Sinica (English Edition): Vol. 79, #5, pp. 593-597 [from abstract] - You, H.-l., Xu, X. & Wang, X.-l., 2003: A New Genus of Psittacosauridae (Dinosauria: Ornithopoda) and the Origin and Early Evolution of Marginocephalian Dinosaurs.
–Acta Geologica Sinica: Vol. 77, #1 - Zhao, X., Cheng, Z., & Xu, X., 1999: The earliest ceratopsian from the Tuchengzi Formation of Liaoning, China.
–Journal of Vertebrate Paleontology: Vol. 19, #4, pp. 681-691 - Zhao, X., Cheng, Z., Xu., X., & Makovicky, P. J., 2006: A New Ceratopsian from the Upper Jurassic Houcheng Formation of Hebei, China.
–Acta Geologica Sinica: Vol 80, #4, pp.467-473 - Zhou, C.-F., Gao, K.-Q., Fox, R. C. & Chen, S.-H., 2006: A new species of Psittacosaurus (Dinosauria: Ceratopsia) from the Early Cretaceous Yixian Formation, Liaoning, China.
–Palaeoworld: Vol. 15, #1, pp. 100-114 [doi: 10.1016/j.palwor.2005.11.001] - Xu, X., Forster, C. A., Clark, J. M. & Mo, J., 2006: A basal ceratopsian with transitional features from the Late Jurassic of northwestern China.
–Proceedings of the Royal Society of London: Biological series: Early Online Publishing, pp. 1-6 [DOI:10.1098/rspb.2006.3566] - Xu, X., Makovicky, P. J., Wang, X.-l., Norell, M. A. & You, H.-l., 2002: A ceratopsian dinosaur from China and the early evolution of Ceratopsia.
–Nature: Vol. 416, pp. 314-317 - Xu, X., Wang, K., Zhao, X., Sullivan, C. & Chen, S., 2010: A New Leptoceratopsid (Ornithischia: Ceratopsia) from the Upper Cretaceous of Shandong, China and Its Implications for Neoceratopsian Evolution.
–Plos One: Vol. 5, #11, art.nr. e13835 [doi: 10.1371/journal.pone.0013835]